Hydrophorinae (Diptera: Dolichopodidae) with description of
Cemocarus stuckenbergi sp.n.
Igor Ya. GRICHANOV
Grichanov, I.Ya. A brief review of the Afrotropical fauna of the
subfamily Hydrophorinae (Diptera: Dolichopodidae) with description of
Cemocarus stuckenbergi sp.n.
Cemocarus stuckenbergi sp.n. from South Africa is described. New
records, keys to the African hydrophorine genera and to recognizable
Afrotropical species of Hydrophorus are given.
I.Ya. Grichanov, All-Russian Institute of Plant Protection,
Podbelskogo 3, St.Petersburg-Pushkin, 189620, Russia.
Key words: Diptera, Dolichopodidae, Hydrophorinae, Cemocarus,
Hydrophorus, Orthoceratium, Tropical and South Africa.
The world fauna of the subfamily Hydrophorinae numbers more than
30 genera, of which 6 genera occur in the Afrotropical Region. The
Palearctic fauna of the subfamily was recently revised by Negrobov
(1977-1979), Nearctic fauna by Hurley (1985), and Oriental fauna
(partly) by Meuffels & Grootaert (1984) and Grootaert & Meuffels
(1988, 1993). The last authors described Cemocarus, a new monotypic
genus of Hydrophorinae from South Africa. Rampini & Munari (1986) gave
a key to known Afrotropical species of the genus Aphrosylus Haliday.
Dyte (1967) and Negrobov e.a. (1987) reviewed known Afrotropical
species of the genus Liancalus Loew. Grichanov (in litt.) gave a list
and a key to known Afrotropical and Palearctic species of the genus
Thinophilus Wahlberg. The first record of the genus Orthoceratium from
Afrotropics is given in this paper. Several other North African genera
can be found in Africa south of Sahara. Catalogue of the Afrotropical
genera and key to all African genera are represented below.
Treating unidentified material from the collections of the Natural
History Museum, London (NHML), the Hungarian Natural History Museum
(HNHM), and Lund University (Lund), I found many interest hydrophorine
species. In this paper descriptions of Cemocarus stuckenbergi sp.n.
from South Africa and new records for known African species are given.
Catalogue and key to species of Hydrophorus Fallen are also
The most interest distribution of species are as follows:
Hydrophorus praecox Lehmann - all zoogeographical regions;
Orthoceratium lacustre (Scopoli) - Europe (except North), Algeria,
Tunisia, Madeira, Tanzania (!); Thinophilus indigenus Becker - Cape
Verde Is., Ethiopia, South Yemen, Nigeria, Benin, Ghana, Zaire,
Tanzania, Madagascar, Angola, Namibia, South Africa, Swaziland;
Palearctic and Oriental Regions; Thinophilus mirandus Becker -
Algeria, Morocco, Tanzania.
Holotype and paratypes of the new species are conserved in the
Natural History Museum (London).
List of Afrotropical genera of Hydrophorinae
Aphrosylus Haliday in Walker, 1851:220
Thinophilus Wahlberg, 1844:37
Hydrophorus Fallen, 1823:2
Liancalus Loew, 1857:22
Orthoceratium Schrank, 1803:55
Cymatopini, new tribe
Cemocarus Meuffels & Grootaert, 1984:152
Key to African genera of Hydrophorinae
1. Labellae hook-shaped in lateral view, with long recurved, generally
- Labellae normal in lateral view, without long protruding
2. Arista apical; fore tibia at apex with distinct erect spinose seta;
male hind basitarsomere simple, without strong seta................
- Arista dorsal; fore tibia without spinose seta at apex; male
hind basitarsomere curved, with strong seta........Teneriffa Becker
3. Acrostichals absent; scutellum with 2 or 4 strong setae...........4
- Acrostichals present, or if absent, then scutellum with 6 setae...6
4. Arista apical (males) or subapical (females); 4 dorsocentrals; wing
somewhat darkened.................................Schoenophilus Mik
- Arista dorsal, other features various.............................5
5. Tibiae usually with strong setae; M1+2 usually curved; at least 5
dorsocentrals, or if 4 dc, then male cercus long, at least half as
long as abdomen................................Thinophilus Wahlberg
- All tibiae without apical setae; R2+3, R4+5, and M1+2 straight and
parallel; 4 dorsocentrals; wing hyaline; male cercus short.........
6. First flagellomere trapezoidal, with subapical arista located in
dorso-apical excavation; 6th and 7th male terga well developed.....
.....................................Cemocarus Meuffels & Grootaert
- First flagellomere rounded, oval or subtriangular, without
dorso-apical excavation; at least 7th male tergum greatly reduced..7
7. Mesonotum with small setae; no more than one pair of dorsocentrals;
acrostichals in two rows; arista subapical....Anahydrophorus Becker
- Mesonotum with several strong dorsocentrals; acrostichals in one
row, rarely absent; arista usually dorsal.........................8
8. Fore femora thickened, ventrally with strong bristles and spines;
first flagellomere with apico-ventral incision...Hydrophorus Fallen
- Fore femora not thickened, without strong ventral bristles or
spines; first flagellomere without incision.......................9
9. Scutellum with 4 setae; hind femora flat; wing veins unmodified
except M1+2 with two right angle bends in males and fair sinuation
in females....................................Orthoceratium Schrank
- Scutellum usually with 6 setae; hind femora cylindric; males
and often females with wing veins variously modified, but M1+2
without double right angle bend......................Liancalus Loew
The genus occurs widely across the whole world including
high-latitude islands and mountains with species confined mostly to
the water surface of various reservoirs. The Afrotropical species of
Hydrophorus are worst described and need redescription of type
material. Based on variable characters, original descriptions do not
comprise detailed drawings of hypopygium. Some species known only from
females should be probably synonymized with other species described by
males, and vice versa. For example, male H. incisicornis is possible
synonym of female H. spinicornis. Most part of known species can not
be reliably identified without studying male hypopygium structures,
especially surstylus. Records of the Palearctic species such as H.
balticus from South Africa should be probably belonged to H.
vaalensis. Several other species are awaiting to be synonymized or
declared Nominae Dubiae in future revisions of type material. The last
key to the African species of the genus (Vanschuytbroeck, 1952) was
based on individually and sexually variable characters. I suspect that
only four following species are really known from the Afrotropical
List of Continental Afrotropical species of Hydrophorus
(for references see Dyte & Smith, 1980)
aureifacies Becker, 1914:124 (male) - Kenya, [?Uganda, ?Tanzania,
?Zaire], ?South Africa
praecox Lehmann, 1822:42 - South Africa, Namibia, Angola, Botswana,
South Arabia, Ethiopia, Mauritania, Gambia, Nigeria, Kenya,
Tanzania, Mauritius, Rodriguez, Aldabra, St. Helena, Cape Verde
Islands, Canary Islands; Palearctic, Nearctic, Neotropical,
Oriental Regions, Australia, New Zealand
= inaequalipes Macquart, 1834:453 (Medeterus)
= schonherri Zetterstedt, 1843:444
= cinerea Perris, 1847:492 (Aphrozeta)
= aestum Loew, 1869:36
= vagus Hutton, 1901:34 (Liancalus)
= breviventris, Becker, 1903:60
= monodi Couturier, 1985:13, n. syn.
spinicornis Loew, 1858:373 (female) - South Africa, Zaire, [?Tanzania]
vaalensis Parent, 1954:226 (female) - South Africa, [?Namibia,
Doubtful species and records
arambourgi Parent, 1938:410 (female) - Kenya
balticus Meigen, 1824:66 (Medeterus) - Palearctic Region, St. Helena,
North and ?South Africa
= chloropus von Roser, 1840:56
chappuisi Parent, 1938:411 - Kenya
congoensis Vanschuytbroeck, 1951:74 - Zaire, Tanzania
hydrophylax Parent, 1939:275 - Uganda, ?Namibia
incisicornis Speiser, 1910:111 (male) - Tanzania
jeanneli Parent, 1938:411 - Kenya, Zaire, Tanzania, ?Namibia
ochraceus Becker, 1914:124 (female) - Kenya, Uganda
variinasutus Vanschuytbroeck, 1951:76 - Zaire
Key to recognizable species of Hydrophorus from Continental Afrotropics
1. Apex of first flagellomere reddish or yellow; wing distinctly
maculated....................spinicornis Loew; incisicornis Speiser
- First flagellomere entirely black; wing monochrome, hyaline or
infumated along veins.............................................2
2. Face entirely greyish-white, without metalic reflection; anterior
tibia apically with sharp dens; wing with yellow veins at
base; dorsal lobe of surstylus with long ventral process...........
- Epistome at least partly shining metallic; clypeus white in males
and yellowish in females; anterior tibia apically without dens;
wing with brown veins; dorsal lobe of surstylus without long
3. Hypandrium as long as surstylus, with rounded apex; epandrium with
narrow setiferous process at base of surstylus; female clypeus
- Hypandrium half as long as surstylus, with conoid apex; epandrium
without such process, with short dens at base of surstylus; female
Hydrophorus praecox Lehmann
Material examined. Male and female, Saudi Arabien, W. Buttiker /
Araida, 20.10.75, Selouly's Farm [NHML]; female, Nigeria: Samaru,
15-22.VI.1970, P.H. Ward, B.M. 1970-604 / Mercury vapour light trap;
female, Tanzania: Matombo, Morogoro reg. / 11.IX.1977, leg. Mahunka
[HNHM]; male & female, Gambia: 3 km NW Central Banjul garden at Wadner
Beach Hotel, 21.II.1977, Loc. No. 1A, At light 19.00 - 21.00, UTM
28PCK257891 / Lund Univ., Syst. Dept., Sweden Gambia/Senegal.
Febr.-March 1977, Cederholm - Danielsson - Larsson - Mirestrom -
Norling - Samuelsson; male, Gambia: oil palm and mangrove vegetation
close to the beach, about 5 km SSW Gunjur. At light 19.00 - 22.00,
22.II.1977, Loc. No. 8, UTM 28PCK05-54 / Lund Univ., Syst. Dept.,
Sweden Gambia/Senegal. Febr.-March 1977, Cederholm - Danielsson -
Larsson - Mirestrom - Norling - Samuelsson; 2 males & 1 female, South
West Africa (W22), Kuiseb river canyon, 22-23.I.1972, Riverside
vegatation / Southern African Exp. B.M. 1972-1; 2 males & 1 female,
South West Africa (W32), Windhoek, Race-Course, 5.II.1972, Low
vegatation / At light / Southern African Exp. B.M. 1972-1; 1 male,
South West Africa (W24), Walvis bay, 25-26.I.1972, Pool edge in dunes
/ Southern African Exp. B.M. 1972-1; 1 male, South West Africa (W32),
Windhoek, environs, 6.II.1972, At light / Southern African Exp. B.M.
1972-1; 7 males & 1 female, S.W. Africa (18), Sossusvlei, Diamond Area
No.2, 20-21.I.1972 / Southern African Exp. B.M. 1972-1; 7 males & 1
female, S.W. Africa (W43), Onguma Fm., 55 mls NW Tsumeb, 17-19.II.1972
/ Southern African Exp. B.M. 1972-1; 20 males & 21 females, S.W.
Africa (25), Swakopmund, 26-30.I.1972 / general sweeping / swept
vegetation around sewage farm settling tanks / Southern African Exp.
B.M. 1972-1; 1 male & 1 female, S.W. Africa (11), Aar Farm, 25 mls ESE
Aus, 15-17.I.1972 / Southern African Exp. B.M. 1972-1; 1 male, S.W.
Africa (W36), Otjikoko Sud Fm., 33 mls ENE Omaruru, 10-13.II.1972 /
Southern African Exp. B.M. 1972-1; 1 male, S.W. Africa (19), Sesriem
Canyon, 3 mls W Sesriem, 21-22.I.1972 / Southern African Exp. B.M.
1972-1; 1 male, S.W. Africa (W37), Otjitambi Fm., 27 mls ESE Kamanjab,
13-15.II.1972 / Southern African Exp. B.M. 1972-1; 1 male, S.W. Africa
(17), Sesriem Sud Fm., Maltanhoe distr., 19-20.I.1972 / general
sweeping / Southern African Exp. B.M. 1972-1; 2 females, S.W. Africa
(23), Homeb, 10 mls ESE Gobabeb, 23-25.I.1972 / Southern African Exp.
B.M. 1972-1; 1 female, S.W. Africa (32), Windhoek, 4-10.II.1972 /
Southern African Exp. B.M. 1972-1; 1 male & 12 females, Angola (A34),
Lobito, 19-20.III.1972 / Southern African Exp. B.M. 1972-1; 4 females,
Botswana (B7), Kuke Pan, 20o59'S, 22o25'E, 14-15.IV.1972 / Southern
African Exp. B.M. 1972-1; 2 females, S. Afr. Transvaal, Zontpan N om
Pretoria, 19.6.1955, G. Rudebeck [Lund]; male, [Russian Far East:]
Yevreiskaya AO, vil. Amurzet, 18.08.1991, Grichanov [Author's Coll.];
female, [Kazakhstan:] Tselinograd Region, Novoishimka, 13.08.1989,
Grichanov [Author's Coll.]; 6 males and 1 female, [South Russia:]
Rostov Region, Azov distr., Port-Katon, 17.05.1994, 25.04.1995,
6.06.1996, Grichanov [Author's Coll.].
Diagnosis and variability. H. praecox is well distinguished by
black antenna, entirely greyish-white face, yellow veins at base of
wing, and sharp dens at apex of anterior tibia (see descriptions of H.
praecox by Takagi, 1967, and Negrobov, 1977). Males have dorsal lobe
of surstylus with long ventral process. Many morphological characters
of this species are widely variable. Dorsocentral setae on mesonotum
can be somewhat weaker or stronger, the face narrower or wider, gena
lower or higher, antenna and arista longer or shorter with various
ratio of articles in individuals. The number and strength of ventral
spines on anterior femora are also variable to some extent. Moreover,
hypopygium structures can vary in some specimens (Figs. 1-5). Examined
males from Gambia, South Arabia, and Rostov Region (Russia) have
ventral lobe of surstylus distinctly shorter than dorsal lobe (see
also figures in Takagi, 1967, and Negrobov, 1977), whereas males from
the Russian Far East, Namibia and Angola have ventral lobe reaching
apex of dorsal lobe. The length of ventral process of dorsal lobe, the
number and length of apical and subapical setae on the process and
lobes of surstylus also vary, especially in individuals of South
African population. Most part of 9 examined males (including 3
specimens from Namibia) have hypandrium with conoid apex (as figured
by Negrobov, 1977), whereas males from Gambia and Angola, and a male
from Swakopmund (Namibia) have hypandrium with rounded apex. Couturier
(1985) described H. monodi from Mauritania having no significant
differences in external morphology from H. praecox except wide
membranous area ("sicatrix") on 5th tergum laterally. Examined males
from Gambia have this sicatrix embracing approximately 1/2 to 2/3
lateral surface of 5th tergum and looking like an excavation in dried
specimens. Despite the remark of Couturier, examined males of H.
praecox from Palearctic and Afrotropical Regions also have the
membranous area on both sides of this tergum, although the size of
sicatrix is greatly variable and usually much smaller (from 1/10 to
1/2 lateral surface). The structure of surstylus in Gambian males is
closest to this in European males. This and all other variable
characters do not correlate with each other. So, I regard H. monodi as
synonym of H. praecox.
Distribution: South Africa, Namibia, Angola, Botswana, South
Arabia, Ethiopia, Mauritania, Gambia, Nigeria, Kenya, Tanzania,
Mauritius, Rodriguez, Aldabra, St. Helena, Cape Verde Islands, Canary
Islands; Palearctic, Nearctic, Neotropical, Oriental Regions,
Australia, New Zealand
Hydrophorus ?aureifacies Becker
Material examined. Male, Kenya: Aberdare Range, 26.X.1934,
B.M.E.Afr.Exp. B.M.1935-203 / Mt. Kinangop, 10000 ft, F.W. Edwards; 3
females, Kenya: Mt. Elgon, II.1935, B.M.E.Afr.Exp. B.M.1935- 203 /
Heath Zone, 10500-11500 ft., F.W. Edwards / Alpine Zone, 12000-13000
ft., F.W. Edwards; 4 males & 9 females, Kenya: Mt. Kenya, N.side,
11000-13500 ft / Heather, stream / swept, 20-22.XII.1980, P.S.
Cranston, B.M. 1981-79.
Diagnosis. Examined specimens reveal nearly the same extent of
variation as this in H. praecox. Characters used in original
descriptions and keys to African species (thickness and colour of head
and body pollination, wing infumation, colour of halter and postocular
cilia, length and strength of spines on anterior legs etc.) are
greatly variable and insufficient for reliable identification of
species. It is quite possible, that the following species described
from Equatorial Africa should be synonymized with H. aureifacies (see
also description of this species in Becker, 1923): H. arambourgi
Parent, H. chappuisi Parent, H. congoensis Vanschuytbroeck, H.
hydrophylax Parent, jeanneli Parent, ochraceus Becker, and
variinasutus Vanschuytbroeck. First flagellomere entirely black;
epistome at least partly shining metallic; clypeus white in males and
rusty-yellow in females; anterior tibia apically without dens; wing
with brown veins, usually infumated; colour of halter varies from
dirty-yellow to mostly blackish; hypandrium as long as surstylus, with
rounded apex; epandrium with narrow setiferous process at base of
surstylus; dorsal lobe of surstylus without long ventral process,
slightly sinuate, with strong dorsal seta at base and short ventral
preapical seta; ventral lobe longer than dorsal one, strongly curved
Distribution. Kenya, [?Uganda, ?Tanzania, ?Zaire], ?South Africa.
Hydrophorus ?vaalensis Parent
Material examined. Male & female, RSA: Cape Prov. 15 km E.
Darling, 150 m, 33o26'S, 18o32'E, 04.X.1994, Loc. 3, leg. R.
Danielsson [Lund]; 2 males & 7 females, S. Africa (S6), C.P.
Silvermine, N.R. Cape penin, 2-3.I.1972. / Southern African Exp. B.M.
1972-1; male & female, S. Rhodesia, Luyanga, 16.XI.1948, J.
Diagnosis. Parent (1954) described H. vaalensis by a female from
South Africa, using variable or insignificant characters (see
diagnosis of H. ?aureifacies in this paper). It is impossible to
distinguish this species from other Afrotropical species except H.
spinicornis, H. incisicornis, and H. praecox by published
descriptions. I belong examined material to H. vaalensis, because this
is the only doubtful species, that was described from South Africa.
Records of H. aureifacies, H. balticus, H. jeanneli, and H.
hydrophylax from South Africa and Namibia should be probably refered
to H. vaalensis. The two recognizable species, H. ?aureifacies and H.
?vaalensis, have no significant differences in external morphology
except structures of hypopygium. Wing of the last species is somewhat
paler; halter usually yellow or brownish; hypandrium half as long as
surstylus, with conoid apex; epandrium without process, with short
dens at base of surstylus; dorsal lobe of surstylus without long
ventral process, slightly sinuate, with strong dorsal seta at base and
short ventral preapical seta; ventral lobe longer than dorsal one,
slightly curved at apex; female face yellowish-grey.
Distribution. South Africa, [?Namibia, ?Zimbabwe].
See Dyte, 1967, Negrobov, 1978, 1979b, and Negrobov e.a., 1987
for generic diagnosis, key and descriptions of known species. Six
species are found in Africa from South to Sub-Saharan Region.
Liancalus adenensis Dyte, 1967
Material examined. Female, Ethiopia: Simien, W. of Derasghie, over
9000 ft., 23.XII.1952 / From chapel in cave in basaltic cliff [printed
label] / from damp rock in hoof of chapel (in basaltic rock)
[hand-written label] / N. Ethiopia: 1952-1953, Hugh Scott, B.M.
Diagnosis. L. adenensis differs from other species by 3 pairs of
scutellar setae, short claws on legs (half as long as apical tarsal
segment), uniseriate acrostichals, strongly curved vein M just before
apex. Examined female differs from this described in Dyte (1967) by
smoky spot above M1+2 before its crossing with m-cu.
Distribution: Yemen, Ethiopia (!)
See Negrobov (1978, 1979) for generic diagnosis and descriptions of
two known species. Here is the first record of the genus from the
Orthoceratium lacustre (Scopoli)
Musca lacustris Scopoli, 1763:343
Musca formosus Haliday, 1832:356
Medeterus viridipes Macquart, 1834:452
Orthoceratium lacustre (Scopoli); Negrobov, 1979:419
Material examined. 2 males & 1 female, Tanzania: Kimboza, Forest
Reserve / 11.IX.1977, leg. Mahunka [HNHM];
Diagnosis. O. lacustre differs from the second known species, O.
sabulosum (Becker), by smaller size, wider face, hyaline wing, and
presence of ventral process on 4th and 5th abdominal segments.
Hypopygium structures are identical to these figured by Negrobov
Distribution: Europe (except North), Algeria, Tunisia, Madeira,
Cemocarus Meuffels & Grootaert
See Meuffels & Grootaert (1984) for generic diagnosis and
redescription of type species. The genus has no analogies in
Afrotropical Hydrophorinae. It is related to widely distributed
tropical genus Cymatopus Kertesz, 1901, and Oriental genus
Thinolestris Grootert & Meuffels, 1988. Only two species are known at
Cemocarus griseatus (Curran)
Aphrosylus griseatus Curran, 1926:403;
Cymatopus capensis Parent, 1939:256;
Cemocarus griseatus (Curran) Meuffels & Grootaert, 1984:153.
Material examined. 6 males & 5 females. S.W. Africa (25),
Swakopmund, 26-30.I.1972 / general sweeping / Southern African Exp.
B.M. 1972-1; female, Van Staadens, Pass. E. Cape Prov., S. Africa,
Stuckenberg, V. 1959 [NHML].
Distribution: South Africa, Namibia (!).
Cemocarus stuckenbergi sp.n.
Holotype, male, Van Staadens, Pass. E. Cape Prov., S. Africa /
Coll. B. & P. Stuckenberg, V. 1959 [NHML].
Paratypes, male & female, the same labels.
Description. Frons, face, occiput, palpus, and proboscis with
bronze-black ground color, grey pollinose. A row of strong black
postocular setae, one pair of strong occipital, vertical, and
postvertical setae present. Ocellar tubercle with one pair of strong
setae and several pairs of short hairs. Ventral postcranium with
sparse black irregular cilia supplementing postocular row. Eyes
haired, emarginated near base of antennae. Face wide, the narrowest
near suture. Ratio of height of epistome to its minimal width to
height of clypeus to its maximal width to height of palpus, 20 : 16 :
19 : 26 : 25. Antenna black, scape vase-like, pedicel with short
setae, first flagellomere as long as heigh, trapezoid, with
dorso-apical excavation bearing short arista, with short hairs. Arista
bisegmented, thick and haired at base, otherwise fine and bare. Length
ratio of scape to pedicel to first flagellomere to arista, 12 : 5 : 15
: 32. Palpus and proboscis with short black hairs.
Thorax bronze-black. Six strong dorsocentral bristles, the last
one is the longest; microscopic acrostichals in one irregular row
extending to the 4th dorsocentral seta. Prothorax with several black
cilia. Scutellum with 2 pairs of strong black setae.
Legs mostly black, without strong setae or long hairs,
trochanters and apices of femora light brown, tarsi dark-brown. Fore
and middle coxae with black hairs anteriorly, hind coxa with one fine
external seta. Fore femora and tibia with a row of short black erect
ventral setulae. Fore tibia with short apico-ventral scale. Fifth
tarsomere of all tarsi enlarged and flattened. Length ratio of fore
coxa to femora to tibia to tarsus (segments from first to fifth), 30 :
54 : 42 : 20 : 8 : 6 : 5 : 7. Same ratio for middle leg, 23 : 56 : 57
: 32 : 11 : 6 : 5 : 8. Same ratio for hind leg, 20 : 72 : 69 : 19 : 19
: 9 : 7 : 9.
Wings darkened, veins brown. Ratio of R1 to wing length, 13 : 29.
Ratio of part of costa between R2+3 and R4+5 to this between R4+5 and
M1, 24 : 24. R4+5 and M1+2 straight, parallel in apical part. Ratio of
cross-vein m-cu to apical part of CuA1, 29 : 16. Lower calipter small,
yellow, with dark cilia. Halters yellow, greatly reduced.
Abdomen bronze-green with short black setae; six full terga
developed; 6th tergum nearly as long as 5th one; 7th tergum symmetric,
nearly 2/3 as long as 6th one; 8th segment approximately half as large
as epandrium. Hypopygium black, grey pollinose. Cercus brown,
gradually narrowed apicad, with short hairs along entire length,
reaching the end of surstylus. Surstylus brown, narrow, with rounded
apex, with several scattered short ventral hairs. Hypandrium narrow,
thick at base.
Female. Similar to male except lacking male secondary sexual
characters; fore femora and tibia without erect setulae.
Length: male body without antennae 4.3 mm, wing-length 4.3 mm,
wing-width 1.4 mm; female body 4.8 mm, wing-length 5.0 mm, wing-width
Distribution: South Africa.
Etymology. The species is named in honour of the South African
dipterologist Dr. B.R. Stuckenberg.
Diagnosis. The new species can be separated from C. griseatus
using the following key:
1. Acrostichals biseriate; m-cu approximately as long as apical part
of CuA1; size about 3 mm......................C. griseatus Curran
- Acrostichals uniseriate; m-cu nearly twice as long as apical part
of CuA1; larger species, 4.3
- 4.8 mm.......C. stuckenbergi sp.n.
I am sincerely grateful to Dr. Brian Pitkin, Dr. Laslo Papp, and
Dr. Roy Danielsson for their kindness in furnishing an opportunity to
study the collections of the Natural History Museum (London), the
Hungarian Natural History Museum
(Budapest), and Lund University.
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Aethiopische Region. Entomol. Mitteilungen, 12 (1): 1-50.
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diptere Dolichopodidae de Mauritanie. Revue Fr. Entomol. (N.S.), 7(1):
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Remark under figures
Figs. 1-5. Variation of hypopygium structures in Hydrophorus
praecox Lehmann. 1-2, hypandrium, lateral view; 3-5, dorsal lobe of
surstylus, ventro-lateral view.
Figs. 6-7. Apex of hypopygium, lateral view. 6, Hydrophorus
?aureifacies Becker; 7, Hydrophorus ?vaalensis Parent.
Fig. 8. Cemocarus stuckenbergi sp.n., hypopygium, lateral view.