LIST OF NEW TAXA OF DOLICHOPODIDAE

By FRED CARL HARMSTON
Last updated: 13 September 2010


abaftanus, Dolichopus, Harmston 1966 EN 77(1):6-8. H male (USNM), USA:
Oregon: Howard Prairie Lake Reservoir, 21 miles east of Ashland, 24 Jul. 1963, (L. S.
Miller). A female, P 21 males, 7 females. CS: Same.
aboriginis, Dolichopus, Harmston and Knowlton 1943 BBES 38(3):102-103. H male
(USNM), USA: Utah: Mountain Home, 21 Jul. 1939, (G. F. Knowlton and F. C.
Harmston). A female, P 4 males. CS: Same.
accidentalis, Dolichopus, Harmston and Knowlton 1941 JKES 14(3):93-95, figs. 2,
8. H male (UKC), USA: Colorado: Spring Creek Pass, 29 Jun. 1937, (C. L. Johnson). CS:
Same.
agalmus, Chrysotus, Harmston and Rapp 1968 JKES 41(2):252. H male (CAS),
USA: Nebraska: Glen, 22 Aug. 1960, (F. C. Harmston and W. F. Rapp, Jr.). CS: Same.
alaskensis, Campsicnemus, Harmston and Miller 1966 PESW 68(2):88. H male
(CAS), USA: Alaska: Sterling Highway: Clam Beach, 3 Aug. 1954, (C. P. Alexander). CS:
Same.
albipodus, Hercostomus, Harmston and Knowlton 1941 CE 73(7):131, figs. 8, 9, 11,
13. H male (USNM), USA: Utah: Kanosh Canyon, 12 Jul. 1940, (G. F. Knowlton and F.
C. Harmston). A female, P 5 males. CS: Same.
alexanderi, Campsicnemus, Harmston and Miller 1966 PESW 68(2):89-90. H male
(CAS), USA: Alaska: Taylor Highway, West Fork Dennisan River: Mile Post 49, 13 Aug.
1954, (C. P. Alexander). P 2 males. CS: Same.
alpinus, Medeterus, Harmston and Knowlton 1941 JKES 14(3):95-96, fig. 5. H male
(UKC), USA: California: Alpine, 9 Jul. 1929, (R. H. Beamer). CS: Medetera walschaertsi
Gosseries.
apicales, Peloropeodes, Harmston and Knowlton 1946 (1945) CE 77(8):139. H male
(USNM), USA: Idaho: Coeur dAlene, 28 Sept. 1941, (F. C. Harmston). CS:
Peloropeodes apicalis Harmston and Knowlton.
appendiculatus, Parasyntormon, Harmston and Knowlton 1943 CE 75(4):63-64. H
male (USNM), USA: Utah: Manila, 17 Jul. 1940, (G. F. Knowlton and F. C. Harmston).
A female, P 3 males. CS: Same.
arizonicus, Dolichopus, Harmston 1951 JKES 24(3):103-104. H male (UKC), USA:
Arizona: Huachuca Mountains: Sunnyside Canyon, 9 Jul. 1940, (D. E. Hardy). A female,
P 5 males, 4 females. CS: Same.
arizonicus, Sympycnus, Harmston 1968 EN 79(1):24-25. H male (CAS), USA:
Arizona: Santa Catalina Mountains, 10 Jun. 1938, (O. Bryant). CS: Calyxochaetus arizonicus
(Harmston).
arnaudi, Medeterus, Harmston 1951 GBN 51(1-2):12-13. H male (CAS), USA:
California: Redwood City, 12 Jun. 1948, (P. H. Arnaud, Jr.). A female, P 35 males, 47
females. CS: Medetera arnaudi Harmston.
bajaensis, Chrysotus, Harmston 1968 EN 79(1):23-24. H male (CAS), Mexico:
Bahia [error for Baja] California: Sierra San Pedro Martir, Rancho Viejo, 7,000 ft, 14 Jun.
1953, (P. H. Arnaud, Jr.). CS: Same.
barri, Pelastoneurus, Harmston 1972 EN 83(6): 153-154. H male (CAS), USA:
California: Inyo County: Tecopa Hot Springs, 16 May 1965, (W. F. Barr). A female, P 22
males, 12 females. CS: Same.
beameri, Dolichopus, Harmston and Knowlton 1941 JKES 14(3):92-93, figs. 4, 6. H
male (UKC), USA: Arizona: Chiricahua Mountains, 8 Jul. 1932, (R. H. Beamer). A
female, P 3 males. CS: Same.
binodatus, Sympycnus, Harmston and Knowlton 1940 AESA 33(2):397, figs. 1-3, 7.
H male (USNM), USA: Utah: Leeds, 13 Sept. 1939, (G. F. Knowlton and F. C. Harmston).
CS: Calyxochaetus binodatus (Harmston and Knowlton).
boharti, Tachytrechus, Harmston 1968 EN 79(1):15-16. H male (UCD), USA:
California: Sierra County: Yuba Pass, 7 Jul. 1964, (R. M. Bohart). CS: Same.
cacheae, Hercostomus, Harmston and Knowlton 1941 CE 73(7):131-132, figs. 3, 10.
H male (USNM), USA: Utah: Avon-Liberty road, approx. 5 miles south of Avon, extreme
south end of Cache Valley, 5 Jul. 1940, (G. F. Knowlton and F. C. Harmston). CS: Same.
californica, Melanderia, Harmston 1972 EN 83(6):155-156. H male (CAS), USA:
California: San Diego, salt marsh along ocean, 10 Aug. 1966, (F. C. Harmston). A female,
P 5 males, 7 females. CS: Same. Melanderia (Wirthia) californica Harmston.
californica, Neurigona, Harmston 1972 EN 83(6):154-155. H male (CAS), USA:
California: Santa Cruz County: Greyhound Rock, 22 May 1952, (P. H. Arnaud, Jr.). CS:
Same.
californicum, Syntormon, Harmston 1951 GBN 51(1-2):15-16. H male (UCB)
[CAS], USA: California: Modoc County: 2 miles east of Canby, 12 Jul. 1947, (R. L.
Usinger). CS: Same.
californicus, Aphrosylus, Harmston 1952 PESW 54(6):292-293. S (9 males, 8
females) (USNM), USA: California: Laguna, 1 Aug. 1932, (J. M. Aldrich). CS:
Paraphrosylus californicus (Harmston).
californicus, Polymedon, Harmston and Knowlton 1943 BBES 38(3):105-106. H
male (CAS), USA: California: Sonora, 8 Sept. 1941, (F. C. and V. H. Harmston). CS:
Tachytrechus californicus (Harmston and Knowlton).
californicus, Sympycnidelphus, Harmston 1968 EN 79(1):23. H male (UCD), USA:
California: Amador County: Fiddletown, 18 Jul. 1961, (L. A. Stange). P male. CS: Same.
californicus, Systenus, Harmston 1968 EN 79(1):16-17. H male (CAS), USA:
California: Marin County: Mill Valley, 25 Jun. 1950, (H. B. Leach) [error for Leech]. CS:
Same.
californicus, Tachytrechus, Harmston and Knowlton 1963 JKES 36(4):234-235. H
male (CAS), USA: California: Trinity County: Trinity River Camp, 17 Jul. 1953, (A. T.
McClay). A female, P 10 males, 16 females. CS: Tachytrechus harmstoni Meuffels and
Grootaert, 1999, a new name.
chaetilamellus, Hercostomus, Harmston and Knowlton 1941 CE 73(7):127-129, fig.
1. H male (UKC), USA: California: Orange County, 14 Jul. 1929, (P. W. Oman). P male.
CS: Same.
chlanoflava, Chrysotus, Harmston and Knowlton 1940 JKES 13(2):59-60, fig. 7. H
male (UMNH), USA: North Dakota: Tower City, M.W.R., 23 Jun. 1906. A female, P 2
males, female. CS: Chrysotus chlanoflavus Harmston and Knowlton.
cinctipus, Campsicnemus, Harmston 1968 EN 79(1):17-18. H male (CAS), Canada:
Northwest Territories: Aklavik, 18 May 1931, (O. Bryant). A female, P 2 males. CS:
Same.
classicus, Parasyntormon, Harmston and Knowlton 1943 CE 75(4):64-65. H male
(USNM), USA: Utah: San Juan County: Bluff, 21 May 1938, (G. F. Knowlton and F. C.
Harmston). P 3 males. CS: Parasyntormon classicum Harmston and Knowlton.
coloradensis, Campsicnemus, Harmston and Miller 1966 PESW 68(2):90. H male
(CAS), USA: Colorado: Ward, 21 Jun. 1962, (F. C. Harmston). A female, P 1 female. CS:
Same.
coloradensis, Hercostomus, Harmston 1952 PESW 54(6):286. H male (USNM),
USA: Colorado: Boulder, [No date], (J. M. Aldrich). P 2 males, 4 females. CS: Same.
coloradensis, Scellus, Harmston and James in Harmston and Knowlton 1942 CE
74(5):82-83, figs. 1, 5-6. H male (USNM), USA: Colorado: near Saguache, 12,000 ft., 5
Aug. 1940, (R. H. Painter). A female, P 5 males, 17 females. CS: Same.
colutis, Rhaphium, Harmston and James in Harmston and Knowlton 1942 CE
74(5):83-85, fig. 9. H male (CSU) [USNM], USA: Colorado: Pingree Park, 9 Jul. 1938,
(M. T. James). A female, P 3 males, 3 females. CS: Rhaphium colute Harmston and
James.
condomina, Argyra, Harmston and Knowlton 1946 (1945) CE 77(8):137-138. H
male (USNM), USA: Utah: Wolf Creek Pass, 15 Aug. 1937, (G. F. Knowlton and F. C.
Harmston). A female, P 22 males, 7 females. CS: Same.
condylus, Teuchophorus, Harmston and Knowlton 1946 AMN 36(3):671-672. H
male (USNM), USA: Indiana: Terre Haute, 12 Sept. 1943, (F. C. Harmston). CS: Same.
consanguineus, Diaphorus, Harmston and Knowlton 1963 JKES 36(4):232-233. H
male (CAS), USA: California: Los Angeles County: Tanbark Flat, 13 Jul. 1950, (J. C.
Hall). CS: Same.
consanguineus, Hercostomus, Harmston 1952 PESW 54(6):287-288. H male
(USNM), USA: Washington: Olympia, 2 Jun. 1895, (Kincaid). P male. CS:
Gymnopternus consanguineus (Harmston).
crassipodus, Asyndetus, Harmston 1968 EN 79(1):20-21. H male (CAS), USA:
California: Kern County: Red Rock Canyon, 18 May 1937, (E. P. Van Duzee). CS: Same.
cryptus, Hercostomus, Harmston and Knowlton 1941 CE 73(7):130, figs. 4-6, 14. H
male (USNM), USA: Utah: near Manila: Sheep Creek, 17 Jul. 1940, (G. F. Knowlton and
F. C. Harmston). A female, P 8 males, 6 females. CS: Same.
dakotensis, Argyra, Harmston and Knowlton 1939 JKES 12(3):85, figs. 7-8. H male
(UAES) [USNM], USA: South Dakota: Pineridge, 8 Jul. 1924, (H. C. Severin). CS: Same
dakotensis, Chrysotus, Harmston 1952 PESW 54(6):290. H male (USNM), USA:
South Dakota: Brookings, [no date], (J. M. Aldrich). P 2 males. CS: Same.
diminucosta, Teuchophorus, Harmston and Knowlton 1942 AESA 35(1):21-22, figs.
7, 13. H male (USNM), USA: Utah: Logan Canyon, 4 Aug. 1940, (G. F. Knowlton and F.
C. Harmston). A female, P 8 males, female. CS: Same.
divigatus, Dolichopus, Harmston 1952 PESW 54(6):284. H male (USNM), USA:
Oregon: Mt. Hood: Mt. Hood Meadows, 13 Jul. 1932, (J. M. Aldrich). CS: Same.
dreisbachi, Hercostomus, Harmston and Knowlton 1945 JKES 18(2):80. H male
(USNM), USA: Michigan: Midland County, 19 Jun. 1941, (R. R. Dreisbach). CS:
=Hercostomus chetifer Walker.
dreisbachi, Hydrophorus, Harmston and Knowlton 1963 JKES 36(4):231-232. H
male (USNM), USA: Michigan: Midland County, 24 May 1951, (R. R. Dreisbach). CS:
Same.
duplicatus, Tachytrechus, Harmston 1972 EN 83:157. H male (CAS), USA: Oregon:
Baker, 25 Jul. 1965, (F. C. Harmston). CS: Same.
facirecedens, Dolichopus, Harmston and Knowlton 1939 JKES 12(3):83-84, figs. 4-
6. H male (SDSU) [USNM], USA: South Dakota: Waubay, 22 Jun. 1936, (H. C. Severin).
A female, P male, female. CS: Same.
factivittatus, Dolichopus, Harmston 1966 EN 77(1):10-12. H male (USNM), USA:
Indiana: Terre Haute, 4 Jul. 1944, (F. C. Harmston). P 3 males. CS: Same.
fernandezensis, Sympycnus, Harmston 1955 RCE 4:37-38. H male (USC), Chile:
Juan Fernandez Islands: Masafuera: Quebrada de las Casas, 19 Jan. 1952, (G. Kuschel). A
female, 18 P males, 53 females. CS: Same.
flagellatus, Hercostomus, Harmston 1952 PESW 54(6):293-294. H male (USNM),
USA: Alabama: Mobile, (collected in hold of a ship containing cargo of bananas from
Panama), 2 Apr. 1924, Mobile, No. 1984. CS: Paraclius flagellatus (Harmston).
floridanus, Sciapus, Harmston 1971 FE 54(1):85-86. H male (FSCA), USA: Florida:
Monroe County: Everglades National Park: Middle Cape Sable, 7 Apr. 1966, (H. V.
Weems). P male. CS: Amblypsilopus floridanus (Harmston).
flutatus, Hercostomus, Harmston and Knowlton 1945 JKES 18(2):79. H male
(USNM), USA: Michigan: Bay County, 24 May 1941, (R. R. Dreisbach). CS: Same.
footei, Dolichopus, Harmston 1966 EN 77(1):4-5. H male (USNM), USA: Idaho:
Priest Lake: Kalispell Bay, 7 Jul. 1959, (B. A. Foote). A female, P 2 males, 2 females. CS:
Same.
fulgerus, Dolichopus, Harmston 1966 EN 77(1):14-15. H male (USNM), Canada:
British Columbia: Alaska Highway: Muncho Lake: Mile Post 455, 28 Jun. 1952, (C. P.
Alexander). A female, P male. CS: Same.
fuscalaris, Neurigona, Harmston 1968 EN 79(1):26-27. H male (CAS), USA:
Arizona: Santa Catalina Mountains, 15 Jun. 1938, (O. Bryant). A female, P 4 males, 4
females. CS: Same.
fuscitibialis, Sympycnus, Harmston and Knowlton 1940 AESA 33(2):400-401, figs.
5, 12, 19, 22, 24. H male (USNM), USA: Utah: Duchesne County: Currant Creek, 29 Jun.
1939, (G. F. Knowlton and F. C. Harmston). CS: =Calyxochaetus cilifemoratus (Van
Duzee).
georgianus, Neurigona, Harmston and Knowlton 1963 JKES 36(4):234. H male
(USNM), USA: Georgia: Atlanta, 6 Jun. 1943, (H. D. Pratt). CS: Same.
hardyi, Dolichopus, Harmston 1951 JKES 24(3):105-106. H male (UKC), USA:
Arizona: Chiricahua Mountains: Rustlers Park, 5 Jul. 1940, (R. H. Beamer). A female, P
3 males, 3 females. CS: Same.
hardyi, Sympycnus, Harmston and Knowlton 1940 AESA 33(2):397-398, figs. 8-9,
18; H male (USNM), USA: Utah: Hooper, 15 Jun. 1937, (D. E. Hardy). CS:
Calyxochaetus hardyi (Harmston and Knowlton).
hendersoni, Parasyntormon, Harmston and Knowlton 1939 EN 50(9):256-257, figs.
2, 5. H male (UAES) [USNM], USA: Utah: Monticello, 28 Jul. 1938, (G. F. Knowlton and
F. C. Harmston). CS: Same.
hirsutitarsis, Dolichopus, Harmston 1952 PESW 54(6):281-282. H male (USNM),
USA: California: Klamath River, near mouth, 11 Jul. 1930, (J. M. Aldrich). A female, P 7
males, 2 females. CS: Same.
idahoensis, Medeterus, Harmston and Knowlton 1943 BBES 38(3):106-107. H male
(CAS), USA: Idaho: Sandpoint, 28 Sept. 1941, (F. C. and V. H. Harmston). CS:
=Medetera crassivenis Curran.
idahona, Argyra, Harmston and Knowlton 1946 (1945) CE 77(8):138-139. H male
(USNM), USA: Idaho: Sandpoint, 28 Sept. 1941, (F. C. Harmston). P male. CS: Same.
indianus, Dolichopus, Harmston and Knowlton 1946 AMN 36(3):672-673. H male
(USNM), USA: Indiana: Indianapolis, 21 Mar. 1943, (F. C. Harmston). P 11 males. CS:
Same.
indianus, Hercostomus, Harmston 1952 PESW 54(6):285. H male (USNM), USA:
Indiana: Lafayette, 4 Jul. 1916, (J. M. Aldrich). P 3 males, 3 females. CS: =Hercostomus
tibialis (Van Duzee).
indianus, Paraclius, Harmston and Knowlton 1946 JKES 19(1):24-25. H male
(USNM), USA: Indiana: Indianapolis, 11 Jun. 1944, (F. C. Harmston). A female, P 8
males, 10 females. CS: Tachytrechus indianus (Harmston and Knowlton).
intrudus, Chrysotus, Harmston 1951 JKES 24(3):107-108. H male (UKC), USA:
Oklahoma: Commerce, 16 May 1940, (R. H. Beamer). A female, P 4 males. CS: Same.
iowaensis, Dolichopus, Harmston and Knowlton 1939 JKES 12(3):84-85, figs. 1-3.
H male (SDSU) [USNM], USA: Iowa: Dickinson County, 24 Jun. 1938, (B. G. Berger).
CS: Same.
isoaristus, Sympycnus, Harmston and Knowlton 1940 AESA 33(2):402-403, figs.
25-28. H male (USNM), USA: Utah: Cedar Breaks, 3 Aug. 1938, (G. F. Knowlton and F.
C. Harmston). P 2 males. CS: Calyxochaetus isoaristus (Harmston and Knowlton).
jamesi, Peloropeodes, Harmston and Knowlton 1939 EN 50(9):257-258, figs. 3-4. H
male (USNM), USA: Utah: Blue Creek, R. R. Station, 30 Mar. 1939, (H. E. Dorst, M. W.
Allen, and F. C. Harmston). A female, P 7 males, 3 females. CS: =Sympycnus latitarsis
Van Duzee.
jaquesi, Dolichopus, Harmston and Knowlton 1939 PESW 41(3):87-88, 3 figs. H
male (USNM), USA: Iowa: Mt. Pleasant, 23 May 1938, (B. G. Berger). A female, P 5
males, 6 females. CS: Same.
kansensis, Chrysotus, Harmston 1952 PESW 54(6):289-290. H male (USNM),
USA: Kansas: Lawrence, [no date], (J. M. Aldrich). P male. CS: Same.
kennedyi, Syntormon, Harmston and Knowlton 1942 PESW 44(2):23-24, figs. 3-4,
7, 11. H male (USNM), USA: Colorado: Cameron Pass, 18 Aug. 1940, (G. F.Knowlton).
P male. CS: Same.
knowltoni, Scellus, Harmston 1939 UASAL 16:71-73, figs. 1-6. H male (UAES)
[USNM], USA: Utah: Monticello, 27 Aug. 1938, (G. F. Knowlton and F. C. Harmston). P
2 males. CS: Same.
kuscheli, Hydrophorus, Harmston 1955 RCE 4:35-36. H male (USC), Chile: Juan
Fernandez Islands: Masatierra: Bahia Cumberland, 25 Feb. 1951, (G. Kuschel). A female,
P 33 males, 43 females. CS: Hydatostega kuscheli (Harmston).
kyphotus, Dolichopus, Harmston 1966 EN 77(1):15-17. H male (USNM), USA:
Alaska: Matanuska Valley, 27 Jun. 1952, (C. O. Berg). CS: Same.
leucosetus, Chrysotus, Harmston 1971 FE 54(1):86-87. H male (FSCA), Cuba: U. S.
Navy Base, Guantanamo Bay, 8-19 Feb. 1965. CS: Same.
longilamellus, Diaphorus, Harmston 1971 FE 54(1):87-88. H male (FSCA), USA:
Georgia: Oatland Island, 31 Aug. 1968, (J. M. Harmston). P 2 males. SC: Same.
longilamellus, Hercostomus, Harmston and Knowlton 1940 PESW 42(9):127, figs.
1, 7. H male (USNM), USA: Utah: Kanosh Canyon, 20 Jun. 1939, (G. F. Knowlton and F.
C. Harmston). CS: Same.
magnicornis, Peloropeodes, Harmston and Rapp 1968 JKES 41(2):248-249. H male
(USNM), USA: Nebraska: Plymouth, 18 Aug. 1959, (F. C. Harmston and W. F. Rapp, Jr.).
P 7 males. CS: Same.
mchughi, Diostracus, Harmston 1966 JKES 39(2):224-225. H male (CAS), USA:
Oregon: Latourele Falls, 2 Aug. 1962, (R. A. McHugh). P 2 males. CS: Same.
mchughi, Tachytrechus, Harmston 1972 EN 83(6):156-157. H male (CAS), USA:
Oregon: Latourele Falls, 2 Aug. 1962, (R. A. McHugh). P male. CS: Same.
melanus, Campsicnemus, Harmston and Knowlton 1942 BBES 37(1):11, fig. 8. H
male (USNM), USA: Utah: Torrey, 20 Aug. 1939, (G. F. Knowlton and F. C. Harmston).
A female, P 5 males, female. CS: Same.
michiganus, Dolichopus, Harmston and Knowlton 1945 JKES 18(2):78. H male
(USNM), USA: Michigan: Saginaw County, 14 Jul. 1940, (R. R. Dreisbach). CS:
=Dolichopus setifer Loew.
milleri, Campsicnemus, Harmston 1966 JKES 39(2):226. H male (CAS), USA:
Oregon: Boi-Cope State Park, 25 Apr. 1963, (L. S. Miller). CS: Same.
milleri, Tachytrechus, Harmston 1966 JKES 39(2):223-224. H male (CAS), USA:
Oregon: Crater Lake, 7 Aug. 1963, (L. S. Miller). A female, P 3 males, 11 females. CS:
Same.
monarchus, Dolichopus, Harmston 1968 EN 79(1):13-15. H male (CAS), Canada:
Northwest Territories: Aklavik, 27 Aug. 1931, (O. Bryant). A female, P 5 males, 6
females. CS: Same.
montanus, Campsicnemus, Harmston and Knowlton 1942 BBES 37(1):14, fig. 6. H
male (USNM), USA: Montana: Gardner, 8 Sept. 1940, (F. C. and V. H. Harmston). CS:
Same.
myklebusti, Syntormon, Harmston and Miller 1966 PESW 68(2):90-91. H male
(CAS), USA: Washington: Ilwaco, 19 Jul. 1960, (R. J. Myklebust). A female, P female.
CS: Syntormon flexibile Becker.
nebraskaense, Psilopiella, Harmston and Rapp 1968 JKES 41(2):251-252. H male
(USNM), USA: Nebraska: Halsey: U. S. National Forest: Bessey Division, 11 Jun. 1959,
(F. C. Harmston and W. F. Rapp, Jr.). A female, P 24 females. CS: Sciapus nebraskaensis
(Harmston and Rapp).
neocryptus, Hercostomus, Harmston and Knowlton 1941 CE 73(7):130-131, fig. 15.
H male (CNC), Canada: Alberta: Lethbridge, 18 Jul. 1923, (H. E. Gray). Afemale, P male.
CS: Same.
neomexicanus, Dolichopus, Harmston 1951 JKES 24(3):104-105. H male (UKC),
USA: New Mexico: Cloudcroft, 28 Jun. 1932, (R. H. Beamer). CS: Same.
nevadensis, Asyndetus, Harmston 1968 EN 79(1):18-20. H male (UCD), USA:
Nevada: 15 miles southwest of Winnemucca, 9 Sept. 1959, (T. R. Haig). CS: Same.
nigripes, Parasyntormon, Harmston and Knowlton 1943 CE 75(4):64. H male
(USNM), USA: Utah: Iron County: Little Salt Lake, 7 May 1939, (G. F. Knowlton and F.
C. Harmston). CS: Same.
nigropleurus, Dolichopus, Harmston 1966 EN 77(1):1-2. H male (USNM), USA:
Colorado: Dillon, 13 Aug. 1958, (F. C. Harmston). P 2 males. CS: Same.
nomadus, Dolichopus, Harmston and Knowlton 1942 AESA 35(1):18-19, figs. 2-4,
11. H male (CNC), Canada: Alberta: Cypress Hills, 10 Aug. 1939, (E. H. Strickland). CS:
Same.
nudus, Chrysotus, Harmston and Knowlton 1963 JKES 36(4):233-234. H male
(CAS), USA: California: Mono County: Sardine Canyon, 8,500 ft., 28 Jun. 1951, (J. W.
MacSwain). A female, P 5 females. CS: Chrysotus harmstoni Meuffels and Grootaert,
1999, a new name.
occidentalis, Chrysotimus, Harmston 1951 JKES 24(3):109. H male (UKC), USA:
Arizona: Huachuca Mountains: Sunnyside Canyon, 9 Jul. 1940, (D. E. Hardy). A female,
P 24 males, 16 females. CS: Same.
orbicularis, Hercostomus, Harmston 1952 PESW 54(6):286-287. H male (USNM),
USA: California: San Bernardino Mountains: Mill Creek Canyon, 9 Jun. 1924, (J. M.
Aldrich). P 3 males, 11 females. CS: Same.
oregonensis, Asyndetus, Harmston 1966 JKES 39(2):225-226. H male (CAS), USA:
Oregon: Ana River road, near Summer Lake, 21 Jun. 1963, (L. S. Miller). A female, P 3
males, 13 females. CS: Same.
oregonensis, Campsicnemus, Harmston and Miller 1966 PESW 68(2):88-89. H male
(CAS), USA: Oregon: Barview State Park, 12 Mar. 1963, (L. S. Miller). A female, P 4
males. CS: Same.
oregonensis, Syntormon, Harmston and Knowlton 1942 PESW 44(2):25-26, figs. 2,
8. H male (USNM), USA: Oregon: Portland, 5 Sept. 1940, (F. C. and V. H. Harmston).
CS: Same.
oregonensis, Systenus, Harmston and Miller 1966 PESW 68(2):91-92. H male
(CAS), USA: Oregon: Wilsonville, 3 Jul. 1963, (K. Goeden). P male. CS: Achalcus oregonensis
(Harmston and Miller).
parenti, Sympycnus, Harmston and Miller 1966 PESW 68(2):93. New name for
Sympycnus cilifemoratus Parent, 1932.
pennarista, Sympycnus, Harmston and Knowlton 1940 AESA 33(2):399-400, figs. 4,
10-11, 23. H male (USNM), USA: Utah: Manila, 5 Sept. 1939, (G. F. Knowlton and F. C.
Harmston). P 2 males. CS: Calyxochaetus pennarista (Harmston and Knowlton).
pictipes, Sympycnus, Harmston and Knowlton 1940 AESA 33(2):401-402, figs. 6,
13, H male (USNM), USA: Utah: Kanosh Canyon, 28 Jul. 1939, (G. F. Knowlton and F.
C. Harmston). P male. CS: Calyxochaetus pictipes (Harmston and Knowlton).
purus, Hercostomus, Harmston and Knowlton 1963 JKES 36(4):237-238. H male
(CAS), USA: California: Sacramento County: Galt, 27 Jul. 1952, (E. I. Schlinger). P male.
CS: Gymnopternus purus (Harmston and Knowlton).
pygidus, Peloropeodes, Harmston and Rapp 1968 JKES 41(2):249-250. H male
(CAS), USA: Nebraska: Mills, 16 Jun. 1959, (F. C. Harmston and W. F. Rapp, Jr.). P 3
males. CS: Same.
refulgens, Dolichopus, Harmston 1939 AESA 32(2):349-350, figs. 1-3. H male
(CNC), Canada: Alberta: Fawcett, 6 Jun. 1934, (E. H. Strickland). CS: =Dolichopus
packardi Van Duzee.
rossi, Rhaphium, Harmston and Knowlton 1940 JKES 13(2):60-61, figs. 1, 3, 5-6, 8.
H male (ILNHS), USA: Illinois: Golconda, 18 Apr. 1914. Afemale, P 14 males, 7 females.
CS: Same.
rotundus, Sympycnus, Harmston and Miller 1966 PESW 68(2):93. New name for
Sympycnus calcaratus Parent, 1932. CS: Same.
schlingeri, Hercostomus, Harmston and Knowlton 1963 JKES 36(4):237. H male
(CAS), USA: California: Glendale, 8 Jun. 1952, (E. I. Schlinger). A female, P 3 males,
female. CS: Gymnopternus schlingeri (Harmston and Knowlton).
scopiferus, Asyndetus, Harmston 1952 PESW 54(6):288-289. H male (USNM),
USA: Oregon: Hood River, 14 Jul. 1932, (J. M. Aldrich). P male. CS: Asyndetus scopifer
Harmston.
scopiventris, Thinophilus, Harmston and Knowlton 1940 PPE 16(3):108-109, figs.
1-3. H male (CAS), USA: California: Newark, 27 Jun. 1939, (G. F. Knowlton). P female.
CS: Same.
severini, Asyndetus, Harmston and Knowlton 1939 AESA 32(2):351-352, figs. 8-10.
H male (UAES) [USNM], USA: South Dakota: Martin, 7 Jul. 1924, (H. C. Severin). CS:
Same.
shastaensis, Dolichopus, Harmston 1966 EN 77(1):5-6. H male (USNM), USA:
California: Shasta County: Cassel, 15 Jul. 1955, (W. H. Lange). A female, P 4 males, 9
females. CS: Same.
silvicolus, Chrysotus, Harmston 1951 GBN 51(1-2):11-12. H male (CAS), USA:
California: Tuolumne County: Pinecrest, 29 Jun.18 Aug. 1948 [labeled July 29, 1948],
(P. H. Arnaud, Jr.). A female, P 26 males, 16 females. CS: Chrysotus silvicola Harmston.
silvicolus, Dolichopus, Harmston 1951 JKES 24(3):106-107. H male (UKC), USA:
New Mexico: Cloudcroft, 27 Jun. 1940, (R. H. Beamer). P male. CS: Dolichopus silvico-
la Harmston.
similis, Argyra, Harmston and Knowlton 1940 JKES 13(2):58-59, figs. 2, 4, 9. H
male (ILNHS), USA: Illinois: Dongola, 11 May 1917. CS: Same.
sinualaris, Dolichopus, Harmston 1966 EN 77(1):8-9. H male (USNM), USA: North
Dakota: Bismarck, 25 Aug. 1961, (F. C. Harmston). CS: Same.
smithae, Dolichopus, Harmston 1966 EN 77(1):12-13. H male (USNM), USA:
Alaska: Mt. McKinley National Park: Mile Post 29, 15 Jul. 1952, (M. E. Smith). Afemale,
P 2 females. CS: Same.
sombrea, Neurigona, Harmston and Knowlton 1945 JKES 18(2):77-78. H male
(USNM), USA: Michigan: Midland County, 6 Jun. 1941, (R. R. Dreisbach). P 2 males.
CS: Nepalomyia sombrea (Harmston and Knowlton).
spatulatus, Sympycnus, Harmston and Knowlton 1940 AESA 33(2):398-399, figs.
14-15, 17. H male (USNM), USA: Utah: Hanksville, 1 Aug. 1939, (G. F. Knowlton and F.
C. Harmston). P 2 males. CS: =Calyxochaetus oreas (Wheeler).
spinitarsis, Asyndetus, Harmston 1951 JKES 24(3):108-109. H male (UKC), USA:
California: Palm City, 19 Jul. 1940, (R. H. Beamer). CS: Same.
sporadicum, Dolichopus, Harmston and Knowlton 1942 AESA 35(1):17-18, figs. 1,
10, 12. H male (UAES) [USNM], USA: Utah: Manila, 17 Jul. 1940, (G. F. Knowlton and
F. C. Harmston). CS: Dolichopus sporadicus Harmston and Knowlton.
squamiciliatus, Thrypticus, Harmston and Rapp 1968 JKES 41(2):250-251. H male
(USNM), USA: Nebraska: Crawford, 22 Aug. 1960, (F. C. Harmston and W. F. Rapp, Jr.).
CS: Same.
squamicilliatus, Dolichopus, Harmston 1966 EN 77(1):2-4. H male (USNM), USA:
Colorado: Red Feather Lake, 19 Jun. 1964, (F. C. Harmston). CS: Same.
stanfordi, Hercostomus, Harmston and Knowlton 1940 PESW 42(9):126-127, figs 2,
8-9. H male (USNM), USA: Utah: Cedar City, 12 Jul. 1939, (G. F. Knowlton and F. C.
Harmston). P 4 males. CS: =Hercostomus occidentalis Cole.
stentorius, Pelastoneurus, Harmston 1971 FE 54(1):88-89. H male (FSCA), USA:
Georgia: Oatland Island, 31 Aug. 1968, (J. M. Harmston). CS: Same.
stricklandi, Dolichopus, Harmston and Knowlton 1939 AESA 32(2):350-351, figs.
4-7. H male (CNC), Canada: Alberta: Gull Lake, 27 Jul. 1932, (E. H. Strickland). CS:
Same.
tahoensis, Tachytrechus, Harmston and Knowlton 1940 CE 72(6):114-115, figs. 2, 5.
H male (USNM), USA: California: Tahoe, 19 Jun. 1927, (J. M. Aldrich). P male, 6
females. CS: Same.
tarsipictis, Dolichopus, Harmston and Knowlton 1963 JKES 36(4):236. H male
(USNM), USA: Oregon: Baker, 23 Jul. 1950, (D. R. Maddock and F. C. Harmston). CS:
Same.
texanus, Sympycnidelphus, Harmston 1968 EN 79(1):21-23. H male (USNM), USA:
Texas: Corpus Christi, 30 Aug. 1966, (F. C. Harmston). CS: Same.
torrida, Neurigona, Harmston 1951 GBN 51(1-2):14-15. H male (CAS), USA:
California: Indio, 1 Apr. 1948, (R. Coleman). CS: Same.
torridus, Hercostomus, Harmston and Knowlton 1941 CE 73(7):129-130, figs. 2, 7,
12. H male (USNM), USA: Utah: Leeds, 22 Jun. 1940, (G. F. Knowlton and F. C.
Harmston). A female, P 10 males, 5 females. CS: =Hercostomus setosus (Van Duzee).
truncatus, Hercostomus, Harmston and Knowlton 1940 PESW 42(9):127-128, figs
5-6. H male (USNM), USA: Utah: Kanosh Canyon, 28 Jul. 1939, (G. F. Knowlton and F.
C. Harmston). P male. CS: Same.
uinta, Neurigona, Harmston and Knowlton 1942 CE 74(5):80, figs. 2, 4, 7. H male
(USNM), USA: Utah: Uintah County: near Whiterocks, 25 Jun. 1940, (G. F. Knowlton
and F. C. Harmston). P 15 males. CS: =Neurigona flava Van Duzee.
uintaensis, Syntormon, Harmston and Knowlton 1940 EN 51(5):130-132, figs. 3-4,
6. H male (USNM), USA: Utah: Altonah, 9 May 1939, (G. F. Knowlton and F. C.
Harmston). A female, P 9 males, 6 females. CS: Same.
utahensis, Asyndetus, Harmston and Knowlton 1942 CE 74(5):85, figs. 8, 10. H
male (USNM), USA: Utah: Price, 1 Jun. 1940, (G. F. Knowlton and F. C. Harmston). A
female, P 3 males. CS: Same.
utahensis, Campsicnemus, Harmston and Knowlton 1942 BBES 37(1):13-14, figs.
3, 9. H male (USNM), USA: Utah: Heber, 29 May 1940, (G. F. Knowlton and F. C.
Harmston). P 6 males. CS: Same.
utahensis, Dolichopus, Harmston and Knowlton 1943 BBES 38(3):101-102. H male
(USNM), USA: Utah: Park City, 11 Jul. 1941, (G. F. Knowlton and F. C. Harmston). A
female. CS: Same.
utahensis, Hercostomus, Harmston and Knowlton 1940 PESW 42(9):125-126, figs.
3-4. H male (USNM), USA: Utah: Moab, 19 Jul. 1939, (G. F. Knowlton and F. C.
Harmston). A female, P 5 males, 3 females. CS: Same.
utahensis, Paraclius, Harmston and Knowlton 1946 JKES 19(1):23-24. H male
(USNM), USA: Utah: St. George, 22 Jun. 1940, (F. C. Harmston). A female, P 10 males,
5 females. CS: Same.
utahensis, Sympycnus, Harmston and Knowlton 1939 EN 50(9):258-259, fig. 1. H
male (UAES) [USNM], USA: Utah: Cedar Breaks, 3 Aug. 1938, (G. F. Knowlton and F.
C. Harmston). P male. CS: Same.
utahensis, Syntormon, Harmston and Knowlton 1942 PESW 44(2):24, figs. 1, 5-6,
10. H male (USNM), USA: Utah: Marysvale, 20 Jun. 1940, (G. F. Knowlton and F. C.
Harmston). CS: Same.
utahensis, Systenus, Harmston and Miller 1966 PESW 68(2):92. H male (CAS),
USA: Utah: Moab, (moist cavity of cottonwood tree), 4 Aug. 1957, (F. C. Harmston). P
male. CS: Achalcus utahensis (Harmston and Miller).
utahensis, Tachytrechus, Harmston and Knowlton 1940 CE 72(6):112-114, figs. 1, 3,
4. H male (USNM), USA: Utah: Torrey, 2 Aug. 1939, (G. F. Knowlton and F. C.
Harmston). A female, P 3 males, 4 females. CS: Same.
utahensis, Teuchophorus, Harmston and Knowlton 1942 AESA 35(1):20-21, figs. 5-
6, 8-9. H male (USNM), USA: Utah: Roosevelt, 19 Jul. 1940, (G. F. Knowlton and F. C.
Harmston). A female, P 8 males. CS: Same.
utahna, Argyra, Harmston 1951 GBN 51(1-2):16-17. H male (CAS), USA: Utah:
Monticello, 7 Jul. 1946, (F. C. Harmston). CS: Same.
valgusa, Neurigona, Harmston and Rapp 1968 JKES 41(2):248. H male (USNM),
USA: Nebraska: Creighton, 23 Jun. 1961, (F. C. Harmston and W. F. Rapp, Jr.). CS: Same.
variegatum, Syntormon, Harmston 1952 PESW 54(6):291-292. H male (USNM),
USA: California: Smith River, 21 Jul. 1932, (J. M. Aldrich). CS: Same.
vegetus, Dolichopus, Harmston 1952 PESW 54(6):283-284. H male (USNM), USA:
Alaska: Mt. McKinley National Park, 1932, (F. W. Morand). P 8 males, 7 females. CS:
Same.
vernaae, Dolichopus Harmston and Knowlton, 1940 EN 51(5):129-130, figs. 1-2, 5.
H male (USNM), USA: Utah: Woodruff, 6 Jun. 1939, (G. F. Knowlton and F. C.
Harmston). A female, P 11 males, 11 females. CS: Same.
vinculatus, Thinophilus Harmston and Rapp 1968 JKES 41(2):253-254. H male
(USNM), USA: Nebraska: Antioch, 22 Aug. 1960 (F. C. Harmston and W. F. Rapp, Jr.).
A female, P 6 males. 3 females. CS: Same.
virens, Parasyntormon, Harmston and Knowlton 1943 CE 75(4)65. H male
(USNM), USA: Utah: Wayne County: Henry Mountains: about 15 miles south of
Hanksville, 6 Jun. 1940 (G. F. Knowlton and F. C. Harmston). A female, P 4 males. CS:
Same.
wasatchensis, Hercostomus, Harmston and Knowlton 1943 BBES 38(3):103-105. H
male (USNM), USA: Utah: Mt. Timpanogos, 6 Jul. 1941, (F. C. and V. H. Harmston). A
female, P male. CS: Same.
wirthi, Aphrosylus, Harmston 1951 GBN 51(1-2):13-14. H male (CAS), USA: California:
San Mateo County: Moss Beach, 21 Mar. 1948 (W. W. Wirth). A female, P 10
males, 13 females. CS: Paraphrosylus wirthi (Harmston).
xanthocal, Chrysotus, Harmston and Knowlton 1940 PPE 16(3):110, fig. 4. H male
(CAS), USA: California: Tracy, 15 Nov. 1939, (F. C. and V. H. Harmston). CS: Same.
zionensis, Neurigona, Harmston and Knowlton 1942 CE 74(5):81-82, fig. 3. H male
(USNM), USA: Utah: Zion Canyon, 13 Jul. 1939, (G. F. Knowlton and F. C. Harmston).
CS: Same.
zygomus, Dolichopus, Harmston 1966 EN 77(1):9-10. H male (USNM), USA: Alaska:
Taylor Highway: West Fork Dennisan River: Mile Post 49, 15 Aug. 1954, (C. P. Alexander).
CS: Same.
*******
See HURLEY, RUNYON AND ARNAUD: FRED CARL HARMSTON: MYIA, Volume 7, No.1, pp.1-39, figs. 1-29 (free access)

ACRONYMS FOR JOURNAL CITATIONS: AESA - Annals of the
Entomological Society of America; AMN - The American Midland Naturalist;
BBES - Bulletin of the Brooklyn Entomological Society; CE - The Canadian
Entomologist; EN - Entomological News; FE - The Florida Entomologist;
GBN - The Great Basin Naturalist; JKES - Journal of the Kansas
Entomological Society; PESW - Proceedings of the Entomological Society of
Washington; PPE - The Pan-Pacific Entomologist; RCE - Revista Chilena de
Entomologa; UASAL - Utah Academy of Sciences, Arts and Letters.

ACRONYMS FOR TYPE DEPOSITORIES OF HOLOTYPES: CAS -
California Academy of Sciences, San Francisco; CNC - Canadian National
Collection, Ottawa; CSU - Colorado State University, Ft. Collins; FSCA -
Florida State Collection of Arthropods, Gainesville; ILNHS - Illinois Naturally
History Survey, Champaign; UAES - Utah Agricultural Experiment Station,
Logan; SDSU - South Dakota State University, Brookings; UCB - University of
28 MYIA, Volume 7, No.1, pp.1-39, figs. 1-29
California, Berkeley; UCD - University of California, Davis; UKC - University
of Kansas, Lawrence; UMNH - Utah Museum of Natural History, Salt Lake
City; USC - Universidad de Chile, Santiago; USNM - National Museum of
Natural History, Washington, D.C., [formerly United States National Museum].


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