Wahlberg (Diptera: Dolichopodidae) with descriptions of new species
Igor Ya. GRICHANOV
Grichanov, I.Ya. Notes on Afrotropical and Palearctic species of
the genus Thinophilus Wahlberg (Diptera: Dolichopodidae) with
descriptions of new species
T. munroi setiscutellatus ssp.n. from Namibia, T. ciliventris
sp.n. from South Africa, Botswana, Angola, and Nigeria are described.
New records for known Afrotropical and Palearctic species are given.
Catalogue and key to species of Thinophilus from both Regions are also
present.
I.Ya. Grichanov, All-Russian Institute of Plant Protection,
Podbelskogo 3, St.Petersburg-Pushkin, 189620, Russia.
Key words: Diptera, Dolichopodidae, Thinophilus, key,
Tropical Africa, Palearctic Region
Introduction
Widely distributed halophilous species of the genus Thinophilus
Wahlberg are confined mostly to arid zones and also to sea coast in
many Regions of the world. Concerning the fauna of the Old World, I
regard that one of the centers of the genus diversity is located at
the border dividing Afrotropical and Palearctic Regions. Totally 40
species are known today from the two Regions. 13 species are recorded
only from Afrotropical Region, 10 species - only from Palearctic
Region, whereas 16 Afrotropical and Palearctic species are located in
Saharan-Arabian Transitional Zone. I conditionally refer to this Zone
the following countries mentioned in the Catalogs (Dyte & Smith,
1980; Negrobov, 1991): Morocco, Algeria, Tunisia, Egypt, Sudan,
Ethiopia, Djibouti, Somalia, and Yemen.
The most interest distribution of species are as follows: T.
indigenus Becker - Mongolia, Iran, Turkey, Algeria, Egypt, Cape Verde
Is., Ethiopia, South Yemen, Nigeria (!), Benin (!), Ghana (!), Zaire,
Tanzania (!), Madagascar, Angola (!), Namibia (!), South Africa (!),
Swaziland (!); Oriental Region; T. mirandus Becker - Algeria, Morocco,
Tanzania (!); T. palpatus Parent - Ethiopia, Nigeria, Angola (!),
Namibia (!), Botswana (!). Several Palearctic species had doubtful
records from Tropical Africa, and some Saharan species were also
recorded from South Europe and/or Central Asia. I would expect that
many species had wider distribution than area known up-to-day. These
data induced me to compile the joint key to all Afrotropical and
Palearctic species of Thinophilus (see below).
All the species described and recorded are referred to the
subgenus Thinophilus s.s. T. prudens Curran, probably, cannot be
united with T. versutus Walker in the subgenus Schoenophilus Mik in
spite of the presence of 4 dorsocentral setae on mesonotum in both
species (compare descriptions in Curran, 1926 and Meuffels &
Grootaert, 1984). On the other hand, T. munroi munroi Curran and T.
munroi setiscutellatus ssp.n. should be possibly separated from other
species in new subgenus. Some of the species are known from females,
others - only from males. Therefore, several species cannot be surely
identified without opposite sex of the same series.
Treating unidentified material from the collections of the
Natural History Museum, London [NHML], the Hungarian Natural History
Museum [HNHM], and Lund University, Sweden [Lund], I found many
interest species. In this paper descriptions of T. munroi setiscutellatus
ssp.n. from Namibia, T. ciliventris sp.n. from South Africa, Botswana,
Angola, and Nigeria, and new records for known African and Palearctic
species are given. Catalogue and key to known species of Thinophilus
from both Regions are also present.
Holotypes and paratypes of the new species are deposited in the
Natural History Museum (London). Some specimens collected by author
are conserved in the collection of Voronezh University, Russia [VU].
List of Afrotropical species of Thinophilus unknown from
Palearctic and Transitional Afro-Palearctic Regions
(for references see Dyte & Smith, 1980)
annulitarsis Parent, 1936:323 - Tanzania
aquaticus Becker, 1914:125 - Kenya, Madagascar
bipunctatus Curran, 1926:27 - South Africa, Zaire
calopus Loew, 1852:659 - Mozambique, Zaire
capensis Curran, 1926:26 - South Africa, Zaire
ciliventris sp.n. - South Africa, Botswana, Angola, Nigeria.
imperialis Curran, 1924:228 (Nematoproctus) - South Africa,
Botswana (!), Congo, Ghana, Nigeria, Zaire
munroi munroi Curran, 1926:20 - South Africa
munroi setiscutellatus ssp.n. - Namibia
prudens Curran, 1926:30 - South Africa, Angola (!), Ghana, Zaire,
Senegal (!)
quadrisetus Parent, 1936:324 - Tanzania, Zaire
rex Curran, 1926:23 - South Africa, Zaire
splendidus Vanschuytbroeck, 1951:80 - Zaire
virgatus Curran, 1926:22 - South Africa
List of Palearctic species of Thinophilus unknown from
Afrotropical and Transitional Afro-Palearctic Regions
(for references see Negrobov, 1991)
bicalcaratus Negrobov, 1971:901 - Uzbekistan, Tadjikistan, Turkmenia
brevicilius Negrobov, 1971:901 - Uzbekistan, Tadjikistan, Kirgizia (!)
longipilus Negrobov, 1971:902 - Russian Far East
neptunus Frey, 1915:78 - Sweden
ornatus Negrobov et Grichanov, 1982:105 - Tadjikistan
pollinosus Loew, 1870:58 - China, Mongolia, Tadjikistan
ruficornis Haliday, 1838:184 (Medeterus) - whole Europe, West Siberia
(Omsk Region, Novosibirsk Region), North Kazakhstan, Kirgizia,
Mongolia, North China
= maculicornis Zetterstedt, 1843:474 (Rhaphium)
setosus Negrobov, 1979:433 - Mongolia
spinitarsis Becker, 1907:315 - China, Iran, Tadjikistan (!), South
Ukraine (Kherson Region)
vanschuytbroecki Negrobov, 1971:902 - Azerbaidjan, Turkmenistan,
Afghanistan
List of Palearctic and Afrotropical species of Thinophilus
found in Transitional Afro-Palearctic Region
(for references see Dyte & Smith, 1980; Negrobov, 1991)
achylleus Mik, 1900:73 - Egypt, Tunisia, Italy, Spain
= albidus Macquart, 1849 (Hydrophorus)
argyropalpis Becker, 1910:25 - Egypt, Tunisia, Algeria, South Arabia,
Iran, Uzbekistan, Turkmenia, Tadjikistan (!), Kazakhstan, South
Russia (Volgograd Region), Ukraine (Odessa Region)
atritarsis Parent, 1929:53 - Sudan
flavipalpis Zetterstedt, 1843:472 (Rhaphium) - Egypt, West Europe
(except North), Ukraine (Odessa Region, Zaporozh'e Region,
Crimea), South Russia (Rostov Region), Caucasus, Kazakhstan,
Kirgizia, Mongolia; ?Mozambique
indigenus Becker, 1902:481 - Mongolia, Iran, Turkey, Algeria,
Egypt, Cape Verde Is., Ethiopia, South Yemen, Nigeria (!), Benin
(!), Ghana (!), Zaire, Tanzania (!), Madagascar, Angola (!),
Namibia (!), South Africa (!), Swaziland (!); India, Nepal,
Malaya, Borneo, Philippines, Taiwan
maculatus Parent, 1929:50 - Sudan
mirandus Becker, 1907:112 - Algeria, Morocco, Tanzania (!)
modestus Becker, 1902:50, 75 - Egypt, ?Austria
ochripalpis Becker, 1910:139 - South Yemen, Somalia
palpatus Parent, 1930:101 - Ethiopia, Nigeria, Angola (!), Namibia
(!), Botswana (!)
promotus Becker, 1910:23 - South Yemen, Djibouti
quadrimaculatus Becker, 1902:49 - Egypt, Tunisia, Algeria,
Tadjikistan, Iran; ?Zaire
setulipalpis Bezzi, 1906:302 - Ethiopia (Eritrea), Zaire, ?Italy
spinulosus Parent, 1929:48 - Sudan, Somalia, Nigeria
tinctus Parent, 1929:51 - Sudan, ?Madagascar, Zaire
(Schoenophilus) versutus Haliday in Walker, 1851:192 - Algeria,
Morocco, Central and South Europe, Ukraine (Lugansk Region)
= maculipennis Strobl, 1899:123
(Pseudacropsilus)
Descriptions and new records
Thinophilus quadrimaculatus Becker
Material examined. Male, South Tadjikistan: Tigrovaya Balka,
tugai, 21.VII.1984, Grichanov [Author's Coll.].
Diagnosis. The species is well recognizable by four black lateral
spots on mesonotum and smoky spot near the end of R2+3 and R4+5 on
wing.
Distribution. Egypt, Tunisia, Algeria, ?Zaire; Tadjikistan.
Thinophilus indigenus Becker
Material examined. 2 males, 1 female, Swaziland: 3 km N Simunye,
26o11' S, 31o57' E, 27.X.1994, loc. 36, leg. R. Danielsson [Lund].
Female, RSA: Cape Prov., Cedarberg, 3 km ESE Kriedowkrans, 32o22' S,
18o59' E, 350 m, 06.X.1994, loc. 10, leg. R. Danielsson [Lund].
Female, Gambia: Bakau, Cape St. Mary at Sun Wing Hotel, swept in veg.
along the beach, 5.XI.1977, UTM 28PCK1991, Loc. 26 / Lund Univ., Syst.
Dept., Sweden Gambia/Senegal. Nov. 1977, Cederholm - Danielsson -
Hammarstedt - Hedquist - Samuelsson. Male & female, Manuara Steppe,
1979.IV.20, leg. Eory-Sipos [HNHM]. 5 males & 3 females, S.W. Africa
(W1): Nr. Onseepkans, Orange R. banks, 8-10.I.1972 / Southern African
Exp. B.M. 1972-1. 2 males, Angola (A11): Bruco, 26.II-2.III.1972 /
Southern African Exp. B.M. 1972-1. 2 females, Angola (A32):
Cachoeiras, 20 mls. SW Gabela, 18-19.III.1972 / Southern African Exp.
B.M. 1972-1. Female, S.W. Africa (W52): Swakop R., 3 mls. S.
Okahandja, 7.IV.1972 / Southern African Exp. B.M. 1972-1. Female, S.
Africa (S24): C.P., Dwyka R., Merwevilk-Koup rd., 2.I.1972 / Southern
African Exp. B.M. 1972-1. Female, Nigeria: Ile-Ife, W State, 15 Aug.
1971, Col. J.T. Medler / DD-46 [NHML]. Female, Tanganyika T.:
Morogoro, 29.I.1917, A. Loveridge / Pres. by Imp. Inst. Ent. Brit.
Mus. 1932-301. Female, Gold Coast: Taimong nr. Accra, 18.III.1922, Dr.
J.W. Scott Macfie / From edge of washing pool / Pres. by Imp. Bur.
Ent. Brit. Mus. 1923-6.
Diagnosis. The species can be recognized by five dark spots on
mesonotum and simple tarsi.
Distribution. Mongolia, Iran, Turkey, Algeria, Egypt, Cape Verde
Is., Ethiopia, South Yemen, Nigeria (!), Benin (!), Ghana (!), Zaire,
Tanzania (!), Madagascar, Angola (!), Namibia (!), South Africa (!),
Swaziland (!); India, Nepal, Malaya, Borneo, Philippines, Taiwan.
Thinophilus rex Curran
Material examined. 2 females, S. Africa: R.E. Turner, Brit. Mus.
1932-145 / Cape Province, Heidelberg dist., Breede River, 6.II.1932.
Diagnosis. T. rex differs from other species with maculated
thorax by hyaline wing, four spots on mesonotum and two spots on
scutellum.
Distribution. South Africa, Zaire.
Thinophilus prudens Curran
(Fig. 5)
Material examined. Male & female, Senegal: in forest 1.5 km NE
Djibelor ca. 6.5 km SW Ziguinchor, 8.III.1977, At light 19.00-21.30,
Loc. No. 23A, UTM 28PCJ575885. / Lund Univ., Syst. Dept., Sweden
Gambia/Senegal. Febr.-March 1977, Cederholm - Danielsson - Larsson -
Mirestrom - Norling - Samuelsson. Male. Angola: Huila District, J.
Balfour-Browne, B.M. 1954-797 / Rio Coroca, 8 mls. N. of Porto
Alexandre, 23. VI.1954, s.l. flying round pools / Stn. No. 290.
Female, Angola (A2): Rocadas, R. Cumene, 19-22.II.1972 / at light /
Southern African Exp. B.M. 1972-1;
Description (male from Angola). Frons bronze-green. Black
postocular setae in upper row on either side present, one black
vertical seta laterally on frons, one pair of ocellar setae, one pair
of occipital setae. Ventral postcranium covered with dirty-white
irregular postocular setae. Face metallic, grey pollinose. Clypeus
nearly twice as wide as high. Antenna short, mostly dark-brown, scape
and pedicel ventrally yellow, first flagellomere rounded, with short
terminal hairs. Arista dorsal, bare, thick at base. Length ratio of
first flagellomere to arista, 7 : 36. Palpus yellow, nearly twice as
high as clypeus, greyish pollinose, with sparse dark hairs mainly
along internal margin. Proboscis black, with white hairs.
Thorax bronze-green, grey pollinose, without dark spots. Pleura
densely whitish pollinose. Four strong dorsocentral bristles with
short hair in front of the first one somewhat laterally; no
acrostichals. Propleura with a few white hairs. Scutellum with 2
strong black setae.
Legs yellow, tarsi brown except base of basitarsomeres. Fore coxa
yellow, with short white hairs. Middle and hind coxae dark except
apex, grey pollinose, with a few short light hairs. Fore femora with
ventral row of very fine hairs, twice as long as diameter of femora.
Fore tibia with fine postero-dorsal seta. Length ratio of fore coxa to
femora to tibia to tarsus (segments from first to fifth) - 30 : 45 :
45 : 17 : 6 : 6 : 6 : 7. Middle femora with a row of ventral hairs in
basal half, as long as diameter of femora. Middle tibia with one
antero-dorsal, one postero-dorsal and row of fine ventral hairs in
apical 2/3, as long as tibia diameter. Length ratio of middle coxa to
femora to tibia to tarsus (segments from first to fifth) - 25 : 60 :
60 : 21 : 12 : 9 : 7 : 8. Hind femora with row of sparse black ventral
setae, longer than half of femora diameter, with one preapical
antero-dorsal seta. Hind tibia with ventral and dorsal rows of hairs,
as long as tibia diameter. Length ratio of hind coxa to femora to
tibia to tarsus (segments first to fifth) - 13 : 70 : 65 : 15 : 15 :
10 : 7 : 9.
Wings hyaline, veins brown. R1 ending at basal 1/3 of wing. Ratio
of parts of costa between R2+3 and R4+5 to those between R4+5 and M1 -
21 : 15. R4+5 and M1+2 straight, parallel in apical half. Ratio of
cross-vein m-cu to apical part of CuA1 - 13 : 20. Lower calipter
yellow, small, with short light hairs. Halters yellow.
Abdomen dark bronze-green, flattened dorso-ventrally,
subtriangular in dorsal view, with short dark setulae. Hypopygium
dark-brown. Cercus yellow, very long, band-like, approximately as long
as 4th and 5th terga combined, white setose along entire length, with
rounded apex; all hairs of equal length except the longest 2 or 3
apical cilia, 1/4 as long as cercus. Surstylus simple, narrow, thin in
apical third, ending at apical 1/3 length of cercus, with one strong
dorsal and one strong ventral preapical cilia, 2 apico-dorsal short
setulae.
Female. Similar to male except lacking male secondary sexual
characters; postero-dorsal seta on anterior tibia indiscernible.
Length: body 2.1 mm, antenna 0.4 mm, wing-length 2.3 mm,
wing-width 0.8 mm.
Diagnosis. This very small species can be recognized by 4
dorsocentral setae on mesonotum and very long male cercus reaching
almost to the base of abdomen. Legs yellow. Base of antenna yellow
ventrally. It can be hardly united with T. (Schoenophilus) versutus
Walker in one subgenus. Male from Angola (see description above)
slightly differs from the description of T. prudens by Curran and from
male collected in Senegal. Cercus of the last male somewhat longer, as
long as 3rd, 4th, and 5th terga, with about 10 apical cilia, nearly
half as long as cercus; surstylus ending in front of the middle of
cercus, with somewhat finer preapical cilia. Female from Senegal
differs from female collected in Angola only by distinctly developed
postero-dorsal seta on fore tibia.
Distribution. South Africa, Angola (!), Zaire, Ghana, Senegal (!).
Thinophilus imperialis (Curran)
Material examined. 7 males & 11 females, Gold Coast: Tamale,
November 1916, J.J. Simpson / Pres. by Imp. Inst. Ent. B.M. 1935-576.
Male, N. Nigeria: Ilorin, Apl. 1912, J.W. Scott Macfie, 1912-429
[NHML]. Male, N. Nigeria: Zaria, Samaru, 17.X.1968 / J.C. Deeming,
m.v. trap [NHML]. 2 females, Nigeria: Ile-Ife, W State, 10815 Aug.
1971, Col. J.T. Medler [NHML]. Female, Botswana (B24): R. Shashe, 20
mls. NW Francistown, 24.IV.1972 / Southern African Exp. B.M. 1972-1.
Diagnosis. T. imperialis differs from other species with long
curved apico-ventral setae on anterior tibia by very long cercus
extending to the base of abdomen.
Distribution. South Africa, Botswana (!), Congo, Ghana, Nigeria,
Zaire.
Thinophilus flavipalpis (Zetterstedt)
Material examined. 1 male & 2 females, Russia: Rostov Region,
Azov distr., Port-Katon, 3 & 9.VI.1996, Grichanov [Author's Coll.].
Male, North Kazakhstan: Tselinograd env., Novoishimka, 17.VII.1989,
Grichanov [Author's Coll.]. 30 males and females, Kirgizia: Issyk-Kul
Lake: Anan'evo, Kuturga, Darkhan, Ottuk, 1978-1979, Grichanov [VU]. 9
males and females, Ukraine: Odessa Region: Berezovka, 12.VIII.1977,
Tiligul'skii Liman, 11.VIII.1977, Bolgrad, 18.VIII.1977, Grichanov
[VU].
Diagnosis. This is a large species with black femora, ornamented
anterior tarsus, relatively simple middle tarsus, and short cercus. It
is closely related to T. ornatus Negrobov et Grichanov.
Distribution. Egypt, West Europe (except North), Ukraine (Odessa
Region, Zaporozh'e Region, Crimea), South Russia (Rostov Region),
Caucasus, Kazakhstan, Kirgizia, Mongolia; ?Mozambique.
Thinophilus brevicilius Negrobov
Material examined. 2 males and 1 female, Kirgizia: Issyk-Kul
Lake: Anan'evo, Ottuk, 1978-1979, Grichanov [VU].
Diagnosis. T. brevicilius is closely related to T. setosus
Negrobov and similar to other species having black femora and
ornamented anterior tarsus, differing from these by simple 3rd and 4th
tarsomeres of anterior tarsus, posterior femora without long ventral
setae, and mostly yellow tibiae.
Distribution. Uzbekistan, Tadjikistan, Kirgizia (!).
Thinophilus mirandus Becker
(Fig. 1)
Material examined. 3 males and 2 females, Tanzania: Matombo,
Morogoro reg. / 11.IX.1977, leg. Mahunka [HNHM].
Description. Frons bronze-green, violet in the middle, slightly
grey-brownish pollinose. A row of black postocular setae ending with
postvertical seta at the top of eye present. One black vertical seta
laterally on frons. Ocellar tubercle with one pair of strong setae and
several hairs. Ventral postcranium covered with dense long white
irregular setae. Face metallic blue-green, slightly pollinose, the
narrowest under antennae. Ratio of height of epistome to its minimal
width to height of clypeus to its maximal width to height of palpus,
21 : 12 : 10 : 20 : 30. Antenna short, entirely yellow-orange, first
flagellomere rounded, with short terminal hairs. Arista dorsal, bare,
brownish, thick at base. Length ratio of scape to pedicel to first
flagellomere to arista - 5 : 5 : 9 : 31. Palpus yellow, white
pollinose, with sparse long black hairs. Proboscis brown, with white
hairs.
Thorax olive-bronze-green, entirely grey-brownish pollinose,
without dark spots. Six dorsocentral bristles, the last one is the
longest; no acrostichals. Prothorax and propleura with a few white
hairs. Scutellum with 2 strong black setae and 2 very small hairs.
Legs yellow, apical segments of tarsi partly darkened. Fore coxa
yellow, white pollinose, with black hairs. Middle and hind coxae
olive-green except apex, grey pollinose, each with one external seta,
middle coxa also with dark hairs. Fore femora with a row of black
poster-ventral hairs in apical half, half as long as diameter of
femora. Fore tibia with 2 rows of somewhat elongated ventral setulae,
and a row of dorsal setae of various length, with 3 long black curved
preapical postero-ventral setae; antero-ventral side bare. First
tarsomere with nearly right angle bend, with ventral excavation at
base; baso-ventral prominence bearing a group of very short spinules;
1st and 2nd tarsomeres with a row of elongated posterior setulae; 3rd
and 4th tarsomeres each with a group of long black dorsal hairs, twice
as long as diameter of tarsomere; 4th tarsomere also with one long
dorsal seta, as long as three last segments of fore tarsus together;
5th tarsomere whitish and flattened; all tarsomeres bare ventrally and
darkened at tip. Length ratio of fore coxa to femora to tibia to
tarsus (segments from first to fifth) - 30 : 48 : 40 : 19 : 9 : 6 : 5
: 8. Middle femora with a few dark fine ventral setae in apical two
thirds, nearly half as long as diameter of femora, and with a few
stronger black posterior preapical setae. Middle tibia with two or
three antero-dorsal, two or three postero-dorsal and one ventral
setae. Middle tarsomeres each with a few strong apical setulae; 4th
and 5th tarsomeres flattened, partly white, ventrally bare. Length
ratio of middle coxa to femora to tibia to tarsus (segments from first
to fifth) - 26 : 53 : 57 : 29 : 10 : 7 : 6 : 10. Hind femora with
poster-dorsal and postero-ventral hairs in apical half, shorter than
half-diameter of femora. Hind tibia with 4 or 5 setae on each of
antero-dorsal, postero-dorsal, antero-ventral and postero-ventral
sides. 5th tarsomere slightly flattened, ventrally bare. Length ratio
of hind coxa to femora to tibia to tarsus (segments first to fifth) -
18 : 66 : 75 : 19 : 15 : 10 : 7 : 9.
Wings hyaline, veins mostly brown, partly yellow in basal half.
R1 short, extending to basal third of wing. Ratio of parts of costa
between R2+3 and R4+5 to those between R4+5 and M1 - 26 : 16. R4+5 and
M1+2 nearly parallel at apex, without distinct curvations. Ratio of
cross-vein m-cu to apical part of CuA1 - 20 : 35. Lower calipter
yellow, with white hairs. Halters yellow.
Abdomen olive-bronze-green, with short dark setae; 6th segment
darker; 8th segment with white pubescence and dark hairs. Hypopygium
dark-brown, white pollinose. Cercus yellow, band-like, with rounded
apex, somehow protruding behind the end of surstylus. Surstylus
brown, stick-shape, densely setose in apical third, with two dorsal
setae in the middle. Two epandrial setae at base of surstylus present.
Hypandrium assymmetric, nearly as wide as cercus, reaching the end of
surstylus.
Female. Similar to male except lacking male secondary sexual
characters, otherwise as follows: ratio of height of epistome to its
minimal width to height of clypeus to its maximal width to height of
palpus, 21 : 17 : 13 : 25 : 30. Femora lacking long setae except
several long postero-dorsal hairs in apical third of hind femora. Fore
tibia with 3 strong antero-dorsal, 3 short postero-dorsal and 3 short
ventral setae. Tarsi simple, but 5th tarsomere of all tarsi slightly
flattened.
Length: body without antennae 3.9 - 5.5 mm, wing-length 3.9 - 5.3
mm, wing-width 1.3 - 1.6 mm.
Diagnosis. T. mirandus is easily recognized by the anterior
basitarsomere having nearly right angle bend. Middle femora with
postero-ventral setae in middle part, at least half as long as femora
diameter. Females can be separated by 6 dorsocentrals, entirely yellow
antenna, black setation on palpus, and maculated wing.
Distribution: Algeria, Morocco, Tanzania (!).
Thinophilus spinitarsis Becker
Material examined. Male, South Tadjikistan: Dusti env.,
13.VII.1991, Grichanov [Author's Coll.].
Diagnosis. T. spinitarsis is closely related to T.
vanschuytbroecki Negrobov. It differs from other species with yellow
femora and ornamented anterior tarsus by fairly curved anterior
basitarsomere, hyaline wing, and long dorsal lobe of surstylus in male
genitalia.
Distribution. China, Iran, Tadjikistan (!), South Ukraine
(Kherson Region).
Thinophilus munroi munroi Curran
Material examined. Female, S. Africa: R.E. Turner, Brit. Mus.
1935-73 / Cape Province, Mossel Bay, Dec. 1934.
Diagnosis. T. munroi munroi strongly differs from all other
species by setation of scutellum having 6 to 8 pairs of short spines
in addition to one pair of long setae. It is closely related to T.
munroi setiscutellatus ssp.n., differing by distinctly annulate
posterior tarsus in both sexes, dark setation on anterior coxa and
femora, and many other characters. Wing vein R1 conspicuously shorter
than in T. munroi setiscutellatus, being also unusually long.
Distribution. South Africa.
Thinophilus munroi setiscutellatus ssp.n.
(Fig. 2)
Holotype, male; paratypes, 7 males and 4 females, S.W. Africa
(25), Swakopmund, 26-30.I.1972 / Southern African Exp. B.M. 1972-1.
Description. Frons and occiput bronze-green, densely
grey-brownish pollinose. 7 to 10 short black postocular setae in
irregular upper row on either side, the last upper seta slightly
stronger. One short black vertical seta laterally on frons. One pair
of ocellar setae. Ventral postcranium covered with dense dirty-white
irregular postocular setae. Face densely grey-yellowish pollinose, the
narrowest under antennae. Ratio of height of epistome to its minimal
width to height of clypeus to its maximal width to height of palpus,
20 : 17 : 11 : 24 : 15. Terminal margin of clypeus covered with short
dense white hairs. Antenna short, orange, first flagellomere brown in
apical third, rounded, with short terminal hairs. Arista dorsal, bare,
thick and brown at base, otherwise fine and whitish. Length ratio of
scape to pedicel to first flagellomere to arista - 4 : 3 : 9 : 35.
Palpus whitish-yellow, greyish-white pollinose, with sparse long white
hairs. Proboscis black, with white hairs.
Thorax olive-bronze-green, entirely grey-brownish pollinose,
without dark spots. Six or seven dorsocentral bristles, the last one
is the longest; no acrostichals. Propleura with a few white hairs.
Scutellum with 3 to 5 short black setae on each side in addition to a
pair of normal long bristles.
Legs yellow, apical segments of fore and middle tarsi and apices
of all tarsomeres of hind tarsus darkened. Fore coxa olive-green,
entirely white pollinose, with short white hairs. Middle and hind
coxae olive-green except apex, grey pollinose, with a few short dark
hairs. Fore femora with a few light ventral hairs at base, half as
long as diameter of femora. Fore tibia with very small, almost
indiscernible setae: two antero-dorsal, two postero-dorsal and one
ventral. Length ratio of fore coxa to femora to tibia to tarsus
(segments from first to fifth) - 30 : 47 : 46 : 19 : 8 : 7 : 5 : 7.
Middle femora bears a group of dark antero-ventral hairs in apical
half, hardly as long as diameter of femora, also with one stronger
preapical anterior seta of the same length, and with a few very short
fine white hairs in basal half. Middle tibia with very small setae:
two antero-dorsal, two postero-dorsal and one postero-ventral. Length
ratio of middle coxa to femora to tibia to tarsus (segments from first
to fifth) - 22 : 54 : 57 : 30 : 10 : 8 : 5 : 7. Hind femora without
long hairs. Hind tibia with short setae: two antero-dorsal, two
postero-dorsal and one ventral. Hind basitarsomere slightly broadened
at apex, with apical group of short dense posterior setulae. Length
ratio of hind coxa to femora to tibia to tarsus (segments first to
fifth) - 15 : 80 : 66 : 25 : 14 : 10 : 6 : 9.
Wings hyaline, veins mostly brown, partly yellow in basal half.
R1 long, extending to the middle of wing. Ratio of parts of costa
between R2+3 and R4+5 to those between R4+5 and M1 - 20 : 14. R4+5 and
M1+2 nearly parallel at apex. M1+2 with indistinct curvation. Ratio of
cross-vein m-cu to apical part of CuA1 - 22 : 47. Lower calipter
yellow, with short yellow hairs. Halters yellow.
Abdomen olive-bronze-green, with short dark setulae. Hypopygium
dark-brown, with white pubescence. Cercus yellow, densely white
setose along entire length, spatulate, flat, band-like with rounded
apex, narrowed distad; all hairs of equal length, approximately as
long as width of cercus. Surstylus brown, bilobate, with thin lobes,
approximately as long as cercus, although protruding behind the end of
cercus; external lobe shorter, flat, with 3 apical and 1 subapical
spinose setae; interior lobe longer, cylindrical, narrowed apicad, with
1 dorsal seta in the middle, 1 apical spinose seta, and several
subapical hairs. Epandrial lobe at base of surstylus present. having
long apical seta. Hypandrium broadened in distal half, as wide as
cercus, narrowed at apex, pointed, reaching the end of external lobe
of surstylus.
Female. Similar to male except lacking male secondary sexual
characters, otherwise as follows: ratio of height of epistome to its
minimal width to height of clypeus to its maximal width to height of
palpus, 25 : 21 : 13 : 29 : 20. Arista whitish. Palpus with short
white hairs. Upper row of 6 or 7 short black postocular setae present,
separate row of 2 or 3 same setae at the top of eye, one short
occipital seta, one vertical seta. Fore and middle femora without long
hairs. Otherwise similar to male.
Length: male body without antennae 4.5 - 4.7 mm, wing-length 4.3
- 5.2 mm, wing-width 1.4 - 1.6 mm.
Distribution: Namibia.
Etymology. The subspecies name reflects the presence of
excessively developed setation on scutellum.
Diagnosis. T. munroi setiscutellatus is an allied subspecies for
T. munroi munroi. It can be separated by weaker developed setation on
scutellum, only pale hairs on anterior coxa and femora, and not so
strongly annulate posterior tarsus as in T. munroi munroi. Females
have the same differences. The species forms a separate group,
possibly, subgenus, having such characters as long vein R1 and
excessively setose scutellum.
Thinophilus ciliventris sp.n.
(Fig. 3)
Holotype, male, S. Africa (S19): R. Magalakwena, 23o26' S, 28o37'
E, 26.IV.1972 / Southern African Exp. B.M. 1972-1.
Paratypes, 2 females, the same labels; 1 male & 3 females, Angola
(A32): Cachoeiras, 20 mls. SW Gabela, 18-19.III.1972 / Southern
African Exp. B.M. 1972-1; 2 females, Botswana (B24): R. Shashe, 20
mls. NW Francistown, 24.IV.1972 / Southern African Exp. B.M. 1972-1;
male, N. Nigeria: Zaria, Samaru, 1.VII.1969 / J.C. Deeming, m.v. trap
[NHML].
Description. Frons blue-violet, slightly pollinose. A row of
short black postocular setae ending with a few postvertical hairs at
the top of eye present; one strong occipital seta; one black vertical
seta laterally on frons. Ocellar tubercle with one pair of strong
setae and several short hairs. Ventral postcranium covered with long
dense white irregular setae. Face violet, slightly pollinose, the
narrowest in the middle of epistome. Ratio of height of epistome to
its minimal width to height of clypeus to its maximal width to height
of palpus, 26 : 17 : 14 : 25 : 33. Antenna short, mostly dark-brown,
all segments ventrally yellow-orange, scape with pale interior spine,
first flagellomere rounded, with short terminal hairs. Arista dorsal,
bare, brown, thick at base, otherwise fine. Length ratio of scape to
pedicel to first flagellomere to arista, 12 : 9 : 13 : 51. Palpus
yellow, white pollinose, with sparse short black hairs. Proboscis
black, with white hairs.
Mesonotum metallic green with copper and blue reflection mostly
grey-brownish pollinose, without dark spots. Pleura olive-green, grey
pollinose. Six or seven dorsocentral bristles, the last one is the
longest; no acrostichals. Propleura and prothorax with long white
hairs. Scutellum with 2 strong black setae and 2 very short hairs.
Legs mostly yellow-orange, apical segments of fore and middle
tarsi darkened, apex of hind femora, extreme base and apex of hind
tibia, and hind tarsus brown. Fore coxa olive-green at base, entirely
white pollinose, with dark hairs in apical half, with shorter white
hairs in basal half. Middle and hind coxae olive-green except apex,
grey pollinose, each with one external seta; middle coxa also with a
few short dark hairs, hind coxa with microscopic light hairs. Fore
femora with a row of black ventral setae, half as long as diameter of
femora, with a few longer poster-ventral setae in apical fourth. Fore
tibia with 3 to 5 dorsal and several simple apical setae, with a row
of ventral setulae, more than half as long as tibia diameter, and a
row of shorter antero-ventral spinules along entire length; anterior
side bare. First tarsomere with ventral row of spinules, half as long
as diameter of tarsomere; other tarsomeres with somewhat elongated
terminal setulae. Length ratio of fore coxa to femora to tibia to
tarsus (segments from first to fifth) - 36 : 60 : 50 : 17 : 8 : 6 : 5
: 7. Middle femora with very short black ventral setae in basal half,
also with one stronger preapical anterior seta. Middle tibia with
three antero-dorsal, two postero-dorsal, one antero-ventral and three
postero-ventral setae of various strength. Last three tarsomeres
slightly flattened, with bare ventral side. Length ratio of middle
coxa to femora to tibia to tarsus (segments from first to fifth) - 29
: 63 : 73 : 42 : 13 : 8 : 6 : 9. Hind femora with ventral and
antero-dorsal rows of seta, shorter than half-diameter of femora. Hind
tibia with four antero-dorsal, four postero-dorsal and three ventral
setae of various strength. Length ratio of hind coxa to femora to
tibia to tarsus (segments first to fifth) - 19 : 95 : 105 : 26 : 19 :
11 : 8 : 9.
Wings practically hyaline, veins mostly brown, partly yellow at
base. R1 short, extending to basal third of wing. Ratio of parts of
costa between R2+3 and R4+5 to those between R4+5 and M1 - 30 : 20.
R4+5 and M1+2 nearly parallel at apex, M1+2 without distinct
curvations. Ratio of cross-vein m-cu to apical part of CuA1 - 27 : 29.
Lower calipter yellow, with short white hairs. Halters yellow.
Abdomen mostly bronze-green with copper reflection, with short
dark setulae; 5th segment with violet reflection; 6th segment entirely
violet; 3rd and 4th sternites with a tuft of numerous long white hairs;
8th segment with white pollination and white pubescence. Hypopygium
black, white pollinose. Cercus light-brown, long, flat, band-like,
pointed on apex, with short dark hairs along entire length and one
somewhat longer cilia on extreme apex, reaching the end of surstylus.
Surstylus brown, broad and flat, mitten-like, with dorso-lateral
horn-like process bearing two small setae, with two cilia near
epandrium, and several scattered short hairs. Hypandrium narrow,
slightly broadened in distal half, reaching the middle of surstylus.
Female. Similar to male except lacking male secondary sexual
characters, otherwise as follows: anterior coxa sometimes imparting
mostly olive-green; fore femora without strong setae; anterior tibia
with 3 or 4 antero-dorsal, 3 or 4 postero-dorsal and 2 or 3 ventral
setae; tarsi simple.
Length: male body without antennae 5.8 - 7.1 mm, wing-length 4.8
- 6.2 mm, wing-width 1.4 - 1.6 mm; female body 5.6 - 6.1 mm, wing 5.5
- 6.0 mm.
Distribution: South Africa, Botswana, Angola, Nigeria.
Etymology. The species name reflects the presence of long cilia
on abdominal venter.
Diagnosis. Judging from the examined material and published
descriptions, none Afrotropical or Palearctic species has abdominal
venter with long white hairs. Only T. bicalcaratus bears 2 bundles of
strong black bristles on the 4th sternite of abdomen. Other diagnostic
characters are as follows: anterior coxa yellow haired in basal half;
posterior femora brown from above in apical third, without long
antero-ventral setae; fore tibia and basitarsomere with ventral row of
very short spinules along entire length; propleura with white
setation; m-cu as long as apical part of CuA1.
Thinophilus palpatus Parent
(Fig. 4)
Material examined. Male, N. Nigeria: Zaria, Samaru, 23.II.1968 /
J.C. Deeming, m.v. trap [NHML]; female, Nigeria: N.W. State, Mokwa,
I.A.R., Mile 1, 8-17.VIII.1970, P.H. Ward, B.M. 1970-604; 2 females,
N. Nigeria: Ilorin, 22.02.1912, Dr. J.V. Scott-Macfie / Pres. by Imp.
Inst. Ent. Brit. Mus. 1931-287; male, S.W. Africa (W1): Nr.
Onseepkans, Orange R. banks, 8-10.I.1972 / Southern African Exp. B.M.
1972-1; female, S.W. Africa: Otjitambi Fm., 27 mls ESE Kamanjab,
13-15.II.1972 / Southern African Exp. B.M. 1972-1; male, S.W. Africa
(W30): Ameib Farm, 31.I-2.II.1972, on vegetation around pools /
Southern African Exp. B.M. 1972-1; 2 females, Angola (A2): Rocadas, R.
Cumene, 19-22.II.1972 / at light / Southern African Exp. B.M. 1972-1;
8 females, Angola (A32): Cachoeiras, 20 mls. SW Gabela, 18-19.III.1972
/ Southern African Exp. B.M. 1972-1; 2 females, Angola (A16): 2 mls.
N. Mocamedes, 29.II.1972 / Southern African Exp. B.M. 1972-1; female,
Angola (A37): 5 mls. NE Negola, 25.III.1972 / Southern African Exp.
B.M. 1972-1; female, Botswana (B7): Kuke Pan, 20o59'S, 22o25'E,
14-15.IV.1972 / Southern African Exp. B.M. 1972-1; female, Botswana
(B16): Maun, 21.IV.1972 / Southern African Exp. B.M. 1972-1;
Diagnosis. This is a comparatively small (3.5 mm) species with
simple yellow legs, narrow face, leaf-like male cercus, differing from
T. ruficornis and related species mainly by hypopygium morphology and
wing veins CuA1/m-cu length ratio.
Distribution. Ethiopia, Nigeria, Angola (!), Namibia (!),
Botswana (!).
Thinophilus ruficornis (Haliday)
Material examined. 3 males & 3 females, South Russia: Rostov
Region, Azov distr., Port-Katon, 3, 9, 18 & 26.06.1996, Grichanov
[Author's Coll.]. 7 males & females, Ukraine: Odessa Region:
Khadzhibeiskii Liman, 28.VIII.1977, Tiligulskii Liman, 19.VII.1976,
Berezovka, 12.VIII.1977, Grichanov [VU]. Male, Russia: Novosibirsk
Region, Krasnozerskoe, 20.VI.1989, Grichanov [Author's Coll.]. 12
males & females, Kirgizia: Issyk-Kul Lake: Anan'evo, Kuturga, Ottuk;
1978-1979, Grichanov [VU].
Diagnosis. This is a comparatively small species with simple
yellow legs, narrow face, short and narrow male cercus, and small
surstylus.
Distribution. Whole Europe, West Siberia (Omsk Region,
Novosibirsk Region), North Kazakhstan, Kirgizia, Mongolia, North
China.
Thinophilus argyropalpis Becker
Material examined. 2 males, South Tadjikistan: Dusti env.,
22 & 24.VI.1985, Grichanov & Shamshev [Author's Coll.].
Diagnosis. T. argyropalpis is small species with simple yellow
legs, closely related to T. vanschuytbroecki Negrobov. It differs from
other species by only two setae on scutellum, silvery-white palpus,
grey pollinose face, and partly yellow antenna.
Distribution. Egypt, Tunisia, South Arabia, Iran, Uzbekistan,
Turkmenia, Tadjikistan (!), Kazakhstan, South Russia (Volgograd
Region), Ukraine (Odessa Region).
Key to Afrotropical and Palearctic species of Thinophilus
1. Mesonotum and/or scutellum with distinct dark lateral spots.......2
- Mesonotum without dark lateral spots..............................8
2. Wing with dark spot near the end of R2+3 and R4+5; anterior spot of
mesonotum nearly as large as notopleura, no prescutellar spot......
.............................................quadrimaculatus Becker
- No spot at wing apex..............................................3
3. Mesonotum with additional spot in front of scutellum..............4
- Mesonotum without spot in front of scutellum......................6
4. Mesonotum with four lateral spots..................indigenus Becker
- Mesonotum with six lateral spots...................................5
5. Anterior basitarsomere with simple setulae, without spinules; small
species (2.75 mm)..................................maculatus Parent
- Anterior basitarsomere with a row of very short spinules...........
.........................................splendidus Vanschuytbroeck
6. Wing distinctly maculated at m-cu and on M1+2 curvation............
.................................................bipunctatus Curran
- Wing hyaline......................................................7
7. 5 dorsocentrals; scutellum with one spot; legs mostly yellow,
anterior femora without ventral comb of setae, middle tibia curved
in the middle....................................setulipalpis Bezzi
- At least 6 dorsocentrals; scutellum with 2 lateral spots; legs
mostly black, anterior femora with ventral comb of black setae,
middle tibia simple......................................rex Curran
8. 4 dorsocentrals; small species (2 - 2.5 mm).......................9
- 5 or 6 dorsocentrals present, front one usually short; size usually
larger than 3 mm.................................................10
9. All male femora with long ventral setae; cercus long, reaching
almost to the base of abdomen; legs yellow...........prudens Curran
- Only middle femora with a row of ventral setae, not longer
than femora diameter; cercus very short, triangular-oval; legs
brownish or greyish-yellow with yellow knees.......................
....................................(Schoenophilus) versutus Walker
10. Pedicel long, with apico-ventral lobe..............promotus Becker
- Pedicel without apico-ventral lobe..............................11
11. Scutellum with 3 to 8 pairs of short spines in addition to one
pair of long setae; 1st and 2nd tarsomeres of hind tarsus annulate
with yellow and black............................................12
- Scutellum with at most two pairs of setae........................13
12. Anterior coxa with black setae and pale hairs; anterior femora
with a few long black ventral hairs in basal third; scutellum with
6 to 8 pairs of spines.........................munroi munroi Curran
- Anterior coxa and femora with pale hairs only; scutellum with 3 to
5 pairs of spines.....................munroi setiscutellatus ssp.n.
13. Anterior four femora at least partly black (females usually
indeterminable); male anterior tibia with 2 or 3 strong curved
apical postero-ventral setae.....................................14
- Anterior four femora yellow, sometimes infuscated from above;
anterior tibia with or without apical setae......................18
14. Anterior femora black from above, other femora mostly
reddish-yellow; male cercus very long, extending to the base of
abdomen........................................imperialis (Curran)
- All femora partly black; cercus much shorter....................15
15. 2nd, 3rd and 4th tarsomeres of anterior tarsus with a group of
black setae, that longer than article diameter...................16
- 3rd and 4th tarsomeres of anterior tarsus without long setae.....17
16. 2nd and 3rd tarsomeres of middle tarsus with long flattened setae.
......................................ornatus Negrobov et Grichanov
- Middle tarsus with simple setae...........flavipalpis (Zetterstedt)
17. Posterior femora with ventral setae, half as long as femora
diameter; tibiae mostly dark.......................setosus Negrobov
- Posterior femora ventrally with short setulae; tibiae mostly yellow
...............................................brevicilius Negrobov
18. Males...........................................................19
- Females.........................................................38
19. Posterior coxa with curved apical internal spine; 5 dorsocentrals
present; scutellum with 2 setae..................ochripalpis Becker
- Posterior coxa without spine.....................................20
20. Anterior basitarsomere with internal excavation or incision at
base; antenna yellow.............................................21
- Anterior basitarsomere without excavation or incision at base....23
21. Anterior basitarsomere with nearly right angle bend; middle femora
with postero-ventral setae in middle part, at least half
as long as femora diameter..........................mirandus Becker
- Anterior basitarsomere fairly curved; middle femora with short
setulae in middle part...........................................22
22. Dorsal lobe of surstylus short; wing near m-cu and on M1+2
curvation distinctly maculated; anterior coxa dark in basal
part......................................vanschuytbroecki Negrobov
- Dorsal lobe of surstylus long; wing practically hyaline; anterior
coxa with dark spot near base....................spinitarsis Becker
23. Anterior and/or middle femora ventrally with hairs and bristles,
nearly as long as femora diameter...............................24
- Anterior and middle femora without long ciliation...............31
24. Palpus with black hairs.........................................25
- Palpus with white hairs.........................................29
25. Anterior basitarsomere with ventral row of short but strong black
spines, at least half as long as article diameter................26
- Anterior basitarsomere without ventral spines, with simple setulae
only.............................................................27
26. Apical half of apical part of M1+2 distinctly arcuate, m-cu 2/3 as
long as apical part of CuA1, length 4 mm..........spinulosus Parent
- M1+2 evenly curved, m-cu as long as apical part of CuA1, length 6
to 7 mm...........................................ciliventris sp.n.
27. Wing with spots at m-cu and on M1+2 curvation; mesonotum with a
black spot in front of scutellum; m-cu as long as apical part of
CuA1, length 2.75 mm...............................maculatus Parent
- Wing and mesonotum without such spots; length 3.5 mm.............28
28. Cercus gradually pointed towards apex; m-cu 2/3 as long as apical
part of CuA1........................................capensis Curran
- Cercus broad in the middle, narrowed towards apex; m-cu as long
as apical part of CuA1..............................palpatus Parent
29. Anterior coxa yellow...........................longipilus Negrobov
- Anterior coxa black.............................................30
30. 5 dorsocentrals present; last tarsomeres black on apex; arista
black; size 4.5 mm.....................................calopus Loew
- 6 or 7 dorsocentrals present; last tarsomeres entirely black;
arista white; size 6 mm.............................virgatus Curran
31. Scutellum with 2 strong and 2 short setae; face nearly twice as
high as wide near suture.........................................32
- Scutellum with 2 setae; face approximately as high as wide near
suture...........................................................36
32. 4th sternite of abdomen with 2 bundles of strong black bristles;
anterior coxa with black hairs; posterior femora with long
antero-ventral setae near apex...............bicalcaratrus Negrobov
- 4th sternite of abdomen without bundles of black bristles; anterior
coxa yellow haired in basal half; posterior femora without long
antero-ventral setae.............................................33
33. 3rd and 4th sternites with a tuft of long white hairs; fore tibia
and basitarsomere with ventral row of very short spinules along
entire length.....................................ciliventris sp.n.
- Abdominal venter without tuft of long hairs......................34
34. Anterior basitarsomere with postero-ventral setae longer than
tarsomere diameter; apical part of CuA1 thrice as long as m-cu.....
....................................................pollinosus Loew
- Anterior basitarsomere without long setae; apical part of CuA1 no
more than twice as long as m-cu..................................35
35. Cercus short and narrow, shorter than surstylus; surstylus small,
dorsal lobe hook-shaped on apex, without long dorsal setae; apical
part of CuA1 approximately twice as long as m-cu...................
...............................................ruficornis (Haliday)
- Cercus broad, leaf-shaped, longer than surstylus; surstylus
narrowed towards apex; apical part of CuA1 approximately as long as
m-cu.................................................palpatus Parent
36. Face shining metallic, slightly pollinose; sutural setae
distinctly developed; abdomen with long hairs......achylleus Becker
- Face silvery or greyish-white pollinose; sutural setae small, 1/3 to
1/4 as long as supraalary setae; abdomen with short hairs........37
37. Palpus yellow; face silvery-white pollinose; antenna mostly dark...
..........................................vanschuytbroecki Negrobov
- Palpus silvery-white; face grey pollinose; antenna distinctly
yellow from below...............................argyropalpis Becker
38. 5 dorsocentrals.................................................39
- At least 6 dorsocentrals present, front one usually short........44
39. Palpus with white hairs.........................................40
- Palpus with black hairs.........................................41
40. Anterior coxa yellow............................quadrisetus Parent
- All coxae black.......................................calopus Loew
41. Antenna entirely yellow.........................ochripalpis Becker
- Antenna partly black............................................42
42. Wing strongly infuscated............................tinctus Parent
- Wing hyaline, at most with spots at m-cu and on M1+2 curvation...43
43. Tarsi entirely black.............................atritarsis Parent
- Tarsi partly light................................aquaticus Becker
44. Antenna entirely yellow.........................................45
- Antenna partly black............................................50
45. Palpus with pale setation.......................................46
- Palpus with black setation......................................48
46. Face shining metallic, practically without pollination; scutellum
with 2 setae.........................................achylleus Mik
- Face pollinose; if face weakly pollinose, than scutellum with 2
strong and 2 small setae........................................47
47. Anterior coxa yellow............................quadrisetus Parent
- All coxae black.......................................calopus Loew
48. Wing without spots; tarsi at least partly black...spinulosus Parent
- Wing maculated with 2 smoky spots on m-cu and M1+2 curvation....49
49. Scutellum with 2 setae; 5 dorsocentrals.........ochripalpis Becker
- Scutellum with 2 strong and 2 small setae; 6 dorsocentrals........
...................................................mirandus Becker
50. Palpus with pale setation.......................................51
- Palpus with black setation......................................55
51. Scutellum with 4 setae..........................quadrisetus Parent
- Scutellum with 2 setae..........................................52
52. Hind tarsomeres annulate with yellow and black; 8 dorsocentrals;
no distinct prothoracic setulae; anterior coxa with exclusively
pale fine hairs................................annulitarsis Parent
- Hind tarsomeres non-annulate....................................53
53. Palpus silvery-white; tarsi black; anterior coxa with pale hairs..
...............................................argyropalpis Becker
- Palpus yellow-orange; tarsi mostly yellow.......................54
54. Anterior coxa with pale hairs............vanschuytbroecki Negrobov
- Anterior coxa with black hairs.....................virgatus Curran
55. Hind femora with long dorsal setae..............................56
- Hind femora without long dorsal setation, at most with single
anterior preapical seta.........................................59
56. Upper postocular setae in two rows.................modestus Becker
- Upper postocular setae in one row...............................57
57. Propleura with 2 or 3 strong black setae and white hairs; hind
femora yellow..................................longipilus Negrobov
- Propleura with white setation...................................58
58. Hind femora brown from above in apical third.....ciliventris sp.n.
- Hind femora entirely yellow........................palpatus Parent
59. Apical part of CuA1 thrice as long as m-cu; posterior coxa mostly
yellow.............................................pollinosus Loew
- Apical part of CuA1 no more than twice as long as m-cu..........60
60. Apical part of CuA1 1/3 longer than m-cu; posterior coxa mostly
dark; tibiae at base and at apex dark..............capensis Curran
- Apical part of CuA1 approximately twice as long as m-cu; posterior
coxa mostly yellow; all tarsi and apex of tibiae dark.............
..............................................ruficornis (Haliday)
Acknowledgements
I am sincerely grateful to Dr. Brian Pitkin, Dr. Laslo Papp, and
Dr. Roy Danielsson for their kindness in furnishing an opportunity to
study the collections of the Natural History Museum (London), the
Hungarian Natural History Museum (Budapest), and Lund University.
References
Curran C.H. 1926. Records of African Dolichopodidae with
descriptions of new species. Rev. Zool. Afr., 14(1): 1-39.
Dyte C.E. & Smith K.G.V. 1980. Family Dolichopodidae. In R.W.
Crosskey (ed.). Catalogue of the Diptera of the Afrotropical Region.
Brit. Mus. (Nat. Hist.), London: 443-463.
Grootaert, P. & Meuffels, H.J.G. 1984. Dolichopodidae (Diptera)
from Papua New Guinea. II. Some new species of the genus Thinophilus
Wahlberg, 1844, from the lowland. Indo-Malayan Zoology, 2: 209-223.
Negrobov O.P. 1991. Dolichopodidae. In Catalogue of Palaearctic
Diptera. V.7. Dolichopodidae-Platypezidae. Akad. kiado. Budapest:
1-291.
Remark under figures
Figs 1-4. Surstylus (=gonopode), lateral view.
1, T. mirandus Becker;
2, T. munroi setiscutellatus ssp.n.;
3, T. ciliventris sp.n.;
4, T. palpatus Parent.
Fig. 5, T. prudens Curran, hypopygium, lateral view.
